Maximum likelihood Chronogram from analysis of 14791 bp of mitochondrial and nuclear DNA depicting relationships within Osteocephalus. Numbers on branches show branch support as percentages: SH-aLRT (before slash) and non-parametric bootstrap support (after). Blue numbers on nodes are estimates of divergence time (in million years). The species name is followed by the number of specimen voucher and the locality.

 
 
  Part of: Venegas PJ, García-Ayachi LA, Toral E, Malqui J, Ron SR (2023) A new species of spiny-backed tree frog, genus Osteocephalus (Anura, Hylidae), from the Yanachaga Chemillén National Park in central Peru. Evolutionary Systematics 7(2): 237-251. https://doi.org/10.3897/evolsyst.7.102360