Corresponding author: Martin Husemann (
Academic editor: Danilo Harms
Types represent the ultimate taxonomic information of a species and hence represent the most important specimens in museums. The entomological collections of the Zoological Museum Hamburg (now part of the Centrum für Naturkunde) hold several thousand primary types of insects. However, despite their importance currently no type database exists and catalogues have not been updated since almost 50 years and are only available in German. Following the publication of our catalogue of
Natural history collections are the main archives of biodiversity. They focus on collecting, maintaining and documenting natural specimens, a crucial task in times of biodiversity decline. The
Despite their importance, the entomological collections in Hamburg are not well documented, nor digitized. Herbert Weidner, a former curator of the collection had published an important series of papers from the 60s until the late 70s, where he documented the collections in much detail (first catalogue -
The
Here, we provide an updated catalogue for the
The table shows all type specimens housed in the collection of the
No. | Genus | Species | Current taxonomy | Type | Count, sex | Author, year | Collector, year | Location | Weidner | Lit. |
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– | H | 1♂ | Ander, 1938 | ? | South-Abbessia, Darassum | M | *1 |
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S | 1♂ 1♀ | Griffini, 1913 | Hamburger S.W. Australische Expedition 1905 | Tamala | Y | *11 |
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– | H | 1♀ | Karny, 1935 | Museum Godeffroy, No. 15838 | Australia, Peak Downs | Y | *18 |
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H | 1♀ | Karny, 1935 | Museum Godeffroy, No. 4590 | Australia, Rockhamptons | Y | *18 |
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SYN of |
H | 1♂ | Griffini, 1913 | 01.06.1905 | Buckland Hill, Stat. 114 near Fremantle | Y | *11 |
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SYN of |
P | 1♀ Larvae | Griffini, 1913 | 01.06.1905 | Buckland Hill, Stat. 114 near Fremantle | Y | *11 | |
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H | 1♂ | Karny, 1935 | F. Sikora, July 1901 | Madagascar, Anevoka | Y | *18 |
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H | 1♂ | Karny, 1935 | H. A. Hedler | Liberia | Y | *18 |
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H | 1♂ | Karny, 1935 | Le Guill, Museum Godeffroy, No. 10138 | Mioko | Y | *18 |
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– | H | 1♀ | Karny, 1935 | Museum Godeffroy, No. 17384 | Ponape | Y | *18 |
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H | 1♂ | Karny, 1935 | ? | Colombia , West South America | Y | *18 |
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– | H | 1♂ | Giglio-Toss, 1895 | J. Bohls | Paraguay | Y | *10 |
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SYN of |
H | 1♂ | Weidner, 1941 | H. Rolle, 14.06.1905 | Africa, Windhuk | Y | *23 |
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SYN of |
P | 1♀ | Weidner, 1941 | K. Wegeleben | Africa, Rehoboth | Y | *23 | |
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– | H | 1♂ | Redtenbacher, 1891 | A. O. Swald ded., 01.04.1890 | Madagaskar, 20 miles north from Tamatave | Y | *20 |
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H | 1♂ | Beier, 1954 | R. Paessler, 14.–21.07.1911 | Costa Rica, Punta Arenas | Y | *3 |
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– | H | 1♀ | Weidner, 1941 | C. v. Erlanger, 12.–18.05.1901 | Africa, Djeroko | Y | *23 |
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– | H | 1♂ | Weidner, 1941 | C. v. Erlanger, 12.04.1901 | Africa, South Abessinien, near mouth of Manelin in Ganale Doria River | Y | *23 |
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– | P | 1♀ | Weidner, 1941 | C. v. Erlanger, 04.-05.01.1901 | Africa, South Abessinien, Dagaja near Mane | Y | *23 | |
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– | P | 1♀ | Redtenbacher, 1891 | ? | Rosario | Y | *20 |
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– | H | 1♂ | Beier, 1949 | H. Rolle, 26.10.1903 | Bolivia, Yungas de la Paz, 1000m | Y | *2 |
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– | P | 1♀ | Beier, 1949 | H. Rolle, 26.10.1903 | Bolivia, Yungas de la Paz, 1000m | Y | *2 | |
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– | P | 1♀ | Beier, 1949 | O. Staudinger, 05.02.1902 | Peru, Marcapata | N | *2 | |
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H | 1♂ | Kaltenbach, 1971 | 01.08–15.07.1959 | Angola, Province Mocamedes, Capolopopo | Y | *12 |
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P | 1♂ | Kaltenbach, 1971 | ? | Republic South-Africa, Port Natal | Y | *12 |
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H | 1♂ | Kaltenbach, 1971 | H. Brauns, 15.12.1898 | Oranje-Freistaat, Bothaville | Y | *12 |
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H | 1♀ | Beier, 1960 | R. Paessler, 26.12.1901 | Guatemala, Champerico | Y | *4 |
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SYN of |
S | 1♂ | Redtenbacher, 1891 | ? | unknown location | Y | *20 |
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L | 1♀ | Krauss, 1890 | R. Greeff | Guinea-Island, St- Thome, Rolas | Y | #3 *19 |
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PL | 1♂ | Krauss, 1890 | R. Greeff | Guinea-Island, St- Thome | Y | #3*19 | |
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H | 1♂ | Beier, 1960 | H. Rolle, 26.10.1903 | Bolivia, Yungas de la Paz, 1000m | Y | *4 |
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P | 1♀ | Beier, 1960 | O. Staudinger, Collection Brunner von Wattenwyl | Peru | Y | *4 |
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– | P | 3♂ 3♀ | Redtenbacher, 1891 | T. Frey | Madagaskar, Nossi-Bé | Y | *20 |
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– | H | 1♂ | Beier, 1960 | 09.09.1926 | Costa Rica, Farm Hamburg at Reventazon | Y | *4 |
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– | P | 1♀ | Beier, 1960 | F. Nevermann, 09.11.1930 | Costa Rica, Parismina river, 3km next to mouth | Y | *4 | |
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– | H | 1♂ | Beier, 1960 | W. Fritsche, 05.10.1913 | Colombia, Pandi, Cundinamarca | Y | *4 |
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– | H | 1♀ | Beier, 1954 | E. Feyer & F. Ohaus, 20.06.1911 | Ecuador, Riobamba-Macas | Y | *3 |
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– | P | 1♀ | Beier, 1954 | J. Michaelis, 22.04.1898 | Brasil, Espirito Santo | Y | *3 |
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– | H | 1♂ | Weidner, 1941 | P. O. Mufindi, 06.07.1932 | Africa, Tanganyika, Kiananga | Y | *23 |
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– | P | 1♀ | Weidner, 1941 | Stierlein, 09.06.1898 | Africa, Uhehe | Y | *23 | |
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– | P | 1♂ | Weidner, 1957 | M. Fontaine, Aug. 1953 | Coll. Mus. Congo, Sankuru, Katako-Kombe | Y | *24 |
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– | P | 1♀ | Weidner, 1957 | Schwetz | Coll. Mus. Congo, Lomami | Y | *24 | |
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SYN of |
H | 1♀ | Weidner, 1941 | A. Bartels, 21.08.1906 | Morocco, Mazagan | Y | *23 |
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– | H | 1♂ | Uvarov, 1936 | H. v. Wissmann, 09.-10.1931 | Arabia, Hadhramaut, Wadi Humum near Mukalla | Y | *22 |
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– | P | 1♀ | Karabag, 1956 | T. Karabag, 29.07.1954 | Turkey, Elazig, Palu, Dabanli | Y | *14 |
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– | H | 1♂ | Beier, 1960 | W. Siegmund, 01.12.1898 | South-America | Y | *4 |
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– | P | 1♀ | Weidner, 1941 | E. Obst, 16.-17.06.1911 | Wakinga-Land between Hohelohe- and Nyarasa-ditch | Y | *23 |
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– | H | 1♂ | Beier, 1960 | F. Nevermann, 12.04.1922 | Costa Rica, plan de Limon, near Las Mercedes, 150–300m, 20–30 km from Atlantic | Y | *4 |
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– | A | 1♀ | Beier, 1960 | F. Nevermann, 22.05.1922 | Costa Rica, plan de Limon, near Las Mercedes, 150–300m, 20–30 from Atlantic | Y | *4 | |
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– | P | 1♂ | Beier, 1960 | F. Nevermann, 12.04.1922 | Costa Rica, plan de Limon, near Las Mercedes, 150–300m, 20–30 km from Atlantic | Y | *4 | |
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– | P | 1♀ | Beier, 1960 | F. Nevermann, 18.02.1922 | Costa Rica, plan de Limon, near Las Mercedes, 150–300m, 20–30 km from Atlantic | Y | *4 | |
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– | P | 1♀ | Beier, 1960 | F. Nevermann, 01.11.1921–30.11.1921 | Costa Rica, plan de Limon, near Las Mercedes, 150–300m, 20–30 km from Atlantic, Farm Hamburg Reventazón 10–30m | Y | *4 | |
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– | P | 1♂ | Beier, 1960 | F. Nevermann, 10.08.1928 | Costa Rica, S. José | Y | *4 | |
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– | P | 1♀ | Beier, 1960 | F. Nevermann, 19.12.1930 | Costa Rica, Salvadora Farm, Eingang Nr. 34 1931 | Y | *4 | |
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– | P | 1♂ 1♀ | Karabag, 1962 | T. Karabag, 04.09.1956 | Turkey, Trabzon, Zigana Daghi, 2500m | Y | *16 |
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– | P | 1♂ 1♀ | Karabag, 1957 | T. Karabag, 02.08.1954 | Turkey, Bingöl daglari, Zarovan yaylasi, 2800m | Y | *15 |
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– | P | 1♂ 1♀ | Karabag, 1962 | May 1952 | Turkey, Thrace, Sinekli | Y | *16 |
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– | P | 1♀ | Beier, 1960 | J. Michaelis, 22.04.1898 | Brasilia, Espirito Santo | Y | *4 |
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– | H | 1♂ | Beier, 1954 | R. Haensch, 30.04.1903 | Ecuador | Y | *3 |
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H | 1♂ | Redtenbacher, 1891 | Museum Godeffroy No. 15041 | Australia, Rockhamptons | Y | *20 |
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SYN of |
L | 1♂ | Charpentier, 1825 | Museum Lüneburg, Coll. Heyer | ? | Y | #3 *7 |
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SYN of |
PL | 1♀ | Charpentier, 1825 | Museum Lüneburg, Coll. Heyer | ? | Y | #2*7 | |
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N | 1♂ | Weidner, 1941 | H. Rolle, Naturkunde Museum Stettin | Africa, Uganda, Entebbe | Y | *23 |
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P | 1♀ | Weidner, 1941 | H. Rolle, Naturkunde Museum Stettin | Africa, Uganda, Entebbe | Y | *23 | |
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– | H | 1♀ | Weidner, 1965 | A. Wagner, 02.10.1905 | Australia, New-South-Wales, Tweed-River | Y | *25 |
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– | H | 1♀ | Beier, 1960 | O. Bürger, 01.01.1898 | Columbia, Villavicentio | Y | *4 |
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– | P | 1♀ | Redtenbacher, 1891 | A. O. Swald , 02.07.1888 | Madagaskar, Nossi-Bé | Y | *20 |
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– | H | 1♀ | Weidner, 1965 | W. Jark, 1931 | Philippins | Y | *25 |
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– | P | 1♀ | Redtenbacher, 1891 | ? | Surinam | Y | *20 |
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– | NA | 1♂ | Chopard, 1957 | F. Sikora, 19.02.1902 | Le Réunion 1901, Plaine des Palmistes | N | #1 *9 |
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– | P | 1♀ | Karabag, 1956 | Ö. K. Güler, 15.07.1952 | Turkey, Adana, Kozan | Y | *14 |
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– | P | 1♂ 2♀ | Karabag, 1964 | T. Karabag, 04.09.1956 | Turkey, Trabzon, Zigana Dagi, 2500m | Y | *17 |
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– | P | 1♀ | Beier, 1960 | H. Rolle, 26.10.1903 | Bolivia, Yungas de la Paz, 1000m | Y | *4 |
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– | P | 1♀ | Kaltenbach, 1971 | H. Brauns, 09.10.1899 | Orange-Freistaat, Bothaville | Y | *12 |
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SYN of |
P | 1♀ | Zeuner, 1931 | Michaelsen, 28.08.1990 | Switzerland, Spiez | Y | *26 |
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SYN of |
P | 1♀ | Zeuner, 1931 | Michaelsen, 20.08.1990 | Switzerland, Beckenried | Y | *26 | |
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– | P | 1♂ 1♀ | Karabag, 1964 | T. Karabag, 27.06.1961 | Turkey, Mozgat, Sorgum, Baglarbasi köyü, Belpinar | Y | *17 |
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– | P | 1♀ | Karabag, 1950 | T. Karabag, 28.06.1949 | Turkey, Ankara, Nallihan | Y | *13 |
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– | P | 1♂ | Karabag, 1950 | T. Karabag, 06.06.1947 | Turkey, Ankara, Orman Ciftligi | Y | *13 |
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– | P | 1♂ | Karabag, 1964 | K.M. Guichard & D. H. Harvey, 23.07.1960 | Turkey, Erzurum, Kop dagi gecidi, 6–7000 ft. | Y | *17 |
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– | P | 1♂ | Karabag, 1962 | T. Karabag, 04.06.1946 | Turkey, Bursa, Karacabey Harasi | Y | *16 |
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– | P | 1♂ | Karabag, 1950 | T. Karabag, 01.06.1946 | Turkey, Anatolia, Bursa, Karacabey, Harasi | Y | *13 |
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– | P | 1♀ | Karabag, 1956 | T. Karabag, 26.06.1954 | Corum, Iskilip | Y | *14 |
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– | S | 1♀ | Redtenbacher, 1891 | E. Versman, 25.09.1886 | East coast of Sumatra, Langkat | Y | *20 |
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– | L | 1♀ | Redtenbacher, 1891 | ? | Siam | Y | #3 *20 |
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SYN of |
P | 1♂ | Redtenbacher, 1891 | Museum Godeffroy | Mioko, Duke of York Island | Y | *20 |
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– | H | 1♀ | Redtenbacher, 1891 | Museum Godeffroy, No. 15037 | Samoa | Y | *20 |
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– | P | 1♀ | Redtenbacher, 1891 | Museum Godeffroy, No. 10095 | Pelew-Islands | Y | *20 |
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– | H | 1♀ | Beier, 1960 | F. Nevermann, 01.12.1926 | Costa-Rica, Plan de Limon near Las Mercedes, Farm Hamburg am Reventazon, 12–30km to Atlantic ocean | Y | *4 |
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– | P | 1♀ | Beier, 1960 | C. Heller, 04.08.1908 | Dutch Guiana, District Cottica | Y | *4 |
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PL | 1♀ | Redtenbacher, 1891 | Museum Godeffroy, No. 2729 | Samoa, Ovalaua | Y | #2 *20 |
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PL | 1♂ | Redtenbacher, 1891 | Museum Godeffroy, No. 2729 | Samoa, Ovalaua | Y | #2 *20 | |
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H | 1♂ | Beier, 1960 | O. Staudinger, 05.02.1902 | Peru, Marcapata | Y | *4 |
The table shows all type specimens that should be in the collection based on the literature and card system, but are currently missing in the collection of the
No. | Genus | Species | Current taxonomy | Type | Count, sex | Author, year | Collector, year | Location | Weidner | Lit. |
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P | 1? | Redtenbacher, 1891 | ? | Brasil, Theresopolis | M | *20 |
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SYN of |
H | 1♂ | Redtenbacher, 1891 | ? | Australia, Peak Downs | M | *20 |
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P | 1? | Redtenbacher, 1891 | ? | Malacca, Perak | M | *20 |
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H | 1♀ | Redtenbacher, 1891 | ? | Uruguay, Montevideo | M | *20 |
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P | ?? | Redtenbacher, 1891 | ? | Argentina, Buenos Aires, Yalapa, Rosario | M | *20 |
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P | ?? | Redtenbacher, 1891 | ? | Mioko | M | *20 |
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SYN of |
P | ?? | Redtenbacher, 1891 | ? | Japan | M | *20 |
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SYN of |
S | ?? | Redtenbacher, 1891 | ? | Brasil, Curitiba, Uruguay, Montevideo | M | *20 |
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P | ?? | Redtenbacher, 1891 | ? | China | M | *20 |
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SYN of |
P | 1♀ | Redtenbacher, 1891 | ? | Singapore | M | *21 |
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SYN of |
P | 1♀ | Redtenbacher, 1891 | ? | Madagascar | M | *20 |
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SYN of |
P | 1♀ | Redtenbacher, 1891 | ? | Argentina, Buenos-Aires | M | *20 |
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H | 1♀ | Redtenbacher, 1891 | ? | Brasil, Lages | M | *20 |
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H | 1♂ | Redtenbacher, 1891 | ? | Australia, Sidney | M | *20 |
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SYN of |
P | ?? | Redtenbacher, 1891 | ? | Surinam | M | *20 |
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P | 1♂ | Redtenbacher, 1891 | ? | Malacca, Perak | M | *20 |
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H | 1♂ | Redtenbacher, 1891 | ? | Mexico | M | *20 |
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S | ?♀ | Redtenbacher, 1891 | ? | Argentina, Buenos-Aires | M | *20 |
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SYN of |
P | 1? | Redtenbacher, 1891 | ? | Brasilia, Theresopolis | M | *20 |
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SYN of |
P | 1? | Redtenbacher, 1891 | ? | West India | M | *20 |
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H | 1♀ | Redtenbacher, 1891 | ? | Australia, Peak Downs | M | *20 |
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P | ?? | Redtenbacher, 1891 | ? | Brasilia, Rio grande do Sul | M | *20 |
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P | ?? | Redtenbacher, 1891 | ? | Uruguay, Montevideo | M | *20 | |
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H | 1♀ | Redtenbacher, 1891 | ? | Australia, Sidney | M | *20 |
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P | 1? | Redtenbacher, 1891 | ? | Australia, North-Queensland | M | *20 |
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- | P | 1? | Redtenbacher, 1891 | ? | ? | M | *20 |
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- | P | 1♀ | Redtenbacher, 1891 | ? | ? | M | *20 |
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S | ?♂ ?♀ | Redtenbacher, 1891 | ? | Australia, Sidney, Rockhampton and New-South-Wales | M | *20 |
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H | 1♀ | Redtenbacher, 1891 | ? | West-Africa, Gaboon | M | *20 |
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H | 1♀ | Redtenbacher, 1891 | ? | Mioko | M | *20 |
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P | 1? | Redtenbacher, 1891 | ? | Australia, Sidney | M | *20 |
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P | ?? | Redtenbacher, 1891 | ? | Yap, Baratonga, Carolinen | M | *20 |
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P | 1? | Redtenbacher, 1891 | ? | Brasilia, Santa Catharina | Y | *20 |
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SYN of |
P | 1? | Redtenbacher, 1891 | ? | Mioko | Y | *20 |
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SYN of |
P | ?? | Redtenbacher, 1891 | ? | Australia, Sidney, Peak Downs, Samo-Islands and Tongatabu | M | *20 |
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- | S | ?♂ ?♀ | Chopard, 1954 | 28.04.1904 | Peru, Cajamarca | M | *8 |
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H | 1♂ | Giglio-Tos, 1895 | J. Bohls | Paraguay | M | *10 |
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- | H | 1♀ | Brunner v. Wattenwyl, 1879 | ? | Australia, Peak Downs | M | *5 |
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- | H | 1♀ | Brunner v. Wattenwyl, 1879 | ? | Australia, Rockhamptons | M | *5 |
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- | S | 1♂ 1♀ | Brunner v. Wattenwyl, 1879 | ? | Australia, Peak Downs and Rockhampton | A | *5 |
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- | S | 1♂ 1♀ | Brunner v. Wattenwyl, 1879 | ? | Australia, Peak Downs | M | *5 |
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- | H | 1♂ | Brunner v. Wattenwyl, 1879 | ? | Australia, Peak Downs | A | *5 |
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- | H | 1♀ | Brunner v. Wattenwyl, 1879 | ? | Australia, Rockhampton | M | *5 |
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- | H | 1♀ | Brunner v. Wattenwyl, 1879 | ? | Australia, Peak Downs | M | *5 |
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- | H | 1♂ | Chopard, 1954 | 09.05.1936 | South-Peru, Esperanza, 900m, near Licht | M | *8 |
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- | H | 1♀ | Brunner v. Wattenwyl, 1895 | ? | Guinea | M | *6 |
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- | S | 12♂♀ | Brunner v. Wattenwyl, 1895 | ? | Perak, Malacca | M | *6 |
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- | H | 1♂ | Beier, 1954 | 06.06.1936 | Costa Rica, Hondura | M | *3 |
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SYN of |
P | 1? | Brunner v. Wattenwyl, 1895 | ? | Venezuela | M | *6 |
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SYN of |
H | 1♀ | Brunner v. Wattenwyl, 1895 | ? | Gabon | M | *6 |
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H | 1♂ | Brunner v. Wattenwyl, 1895 | ? | Mexico, Veracruze, Jalapa | M | *6 |
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- | P | 1♀ | Beier, 1960 | ? | Peru, Sivia | M | *4 |
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- | H | 1♀ | Brunner v. Wattenwyl, 1895 | ? | Gabon | M | *6 |
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- | S | ?? | Brunner v. Wattenwyl, 1895 | ? | Brasilia, Rio de Janeiro | M | *6 |
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H | 1♀ | Brunner v. Wattenwyl, 1895 | ? | Venezuela | M | *6 |
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SYN of |
P | 1? | Brunner v. Wattenwyl, 1895 | ? | Brasilia, Rio de Janeiro | M | *6 |
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- | H | 1♀ | Brunner v. Wattenwyl, 1895 | ? | Cameroon | M | *6 |
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SYN of |
H | 1♀ | Brunner v. Wattenwyl, 1895 | ? | Sumatra, Palenburg | M | *6 |
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- | H | 1♀ | Beier, 1960 | ? | Peru, Sivia, 520m | M | *4 |
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- | H | 1♀ | Beier, 1960 | ? | Peru, Sivia, 520m | M | *4 |
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P | 1? | Brunner v. Wattenwyl, 1895 | ? | Mexico, Jalapa | M | *6 |
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- | H | 1♂ | Beier, 1954 | 15.05.1936 | Peru, Sivia, 520m | M | *3 |
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- | H | 1♂ | Ander, 1938 | Ch. E. Porter | Chile, Pucon at Villarrica lake | M | *1 |
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- | H | 1♀ | Ander, 1938 | ? | Chile, Pucon at Villarrica lake | M | *1 |
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H | 1♂ | Ander, 1938 | Winkler | Borneo, Butik Raja, above 2200m | M | *1 |
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- | H | 1♀ | Chopard, 1954 | 15.05.1936 | South-Peru, jungle | M | *8 |
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- | H | 1♀ | Chopard, 1954 | 12.05.1936 | South-Peru, Sivia, jungle | M | *8 |
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- | H | 1♀ | Chopard, 1954 | 15.05.1936 | Soth-Peru, Sivia, jungle | M | *8 |
Literature: #1
This figure shows
Typical labels of several important taxonomists, who deposited material in the collection:
Two of the 72 species reported by Weidner could not be found during our data acquisition (
In addition to the identified types the collection holds some old specimens which may be types, but require further investigation (e.g. Fig.
Label of the specimen of unknown status (Fig.
The definitions of types are according to the International Commission on Zoological Nomenclature (
Name-bearing types – specimens with a name bearing function (i.e. holotype, syntype, lectotype, neotype)
Holotype – a single specimen on which a new taxon is based in the original publication
Allotype – a specimen of opposite sex of the holotype (no name-bearing function)
Syntype – specimens of a type series that collectively constitute the name-bearing type
Cotype – the term should be avoided; the specimen can be similar to a syntype or a paratype
Lectotype – a specimen that was a syntype, but was designated as unique bearer of a name
Neotype – a name bearing type designated for an existing taxon when no name-bearing type is extant
Paratype – specimens belonging to a type species not being the holotype
Paralectotype – the remaining specimens of a syntype series after the designation of a lectotype
We thank Dr. Holger Braun for many quick and helpful comments. We further thank Josip Skejo, Arne Lehmann and Ionut Stefan Iorgu for their helpful reviews. We are grateful to Thure Dalsgaard for taking some images of specimens. This project was funded by the Orthopterists’ Society, Synthesys and by the Gesellschaft der Freunde und Förderer des Zoologischen Museums Hamburg.