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Research Article
An annotated catalogue of the types of short-horned grasshoppers (Orthoptera, Caelifera) housed in the Zoological Museum Hamburg (ZMH)
expand article infoLara-Sophie Dey‡, Martin Husemann§
‡ Martin-Luther University, Halle, Germany
§ University of Hamburg, Hamburg, Germany

Corresponding author: Martin Husemann ( martin.husemann@uni-hamburg.de )

Academic editor: Danilo Harms
© 2018 Lara-Sophie Dey, Martin Husemann.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation: Dey L-S, Husemann M (2018) An annotated catalogue of the types of short-horned grasshoppers (Orthoptera, Caelifera) housed in the Zoological Museum Hamburg (ZMH). Evolutionary Systematics 2: 21-30. https://doi.org/10.3897/evolsyst.2.22127
ZooBank: urn:lsid:zoobank.org:pub:9C76A3B3-5F8C-4B6A-8A32-AD2FC3A07E24
Open Access

Abstract

Zoological collections remain the main archive of animal life on earth, especially in times of mass extinctions. The most important animals of these collections are the type specimens, which represent the original reference for a species. While a full digitization of most museum collections currently remains utopic, lists of types are an essential resource for researchers to perform taxonomic revisions. Here, we provide an updated type catalogue of the short-horned grasshoppers (Caelifera) housed in the Zoological Museum of Hamburg (ZMH), now part of the Center for Natural History Hamburg (CeNak). This collection comprises 209 type specimens belonging to 128 species: 65 holotypes, three lectotypes, one syntype and a large number of paratypes. We also list topotypes of two species for reasons of completeness but note that they have no formal status. Overall, types of 13 species were added to the collection since the last catalogues in 1968 and 1977 and several corrections to the last catalogues were made. We provide a full bibliography for these species, aiming to provide a comprehensive taxonomic guide to this collection.

Key Words

biodiversity, CeNak, Godeffroy, grasshoppers, museum, type catalogue, ZMH

Introduction

With 209 type specimens of the suborder Caelifera and more than 100 species of Ensifera (Weidner 1966, new type catalogue in preparation) the collection of the Zoological Museum Hamburg (ZMH), now part of the Center of Natural History Hamburg (CeNak), is among the most important collections in Germany. The collection is currently not digitally available, but the former curator Herbert Weidner had presented a first list including types of 116 species and a second publication with four additions (Weidner and Wagner 1968, Weidner 1977; some of these were missing in the collection, when Weidner wrote his catalogue; further, the status of some species was unclear back then). As part of the DORSA project (Ingrisch et al. 2004), the types of the collection were examined and photographed in 2003; these pictures are available from the DORSA website (https://www.dorsa.de/) and have also been implemented in the Orthoptera Species File (Cigliano et al. 2017). However, since the last update of the type catalogue 40 years have passed, rendering a new list necessary. Hence, we present a new and updated list of type specimens of the suborder Caelifera housed in the ZMH with updated nomenclature, and a bibliography of the original descriptions of all species.

The ZMHOrthoptera collection is of special value as many specialists have worked in the museum and deposited material: the museum houses the type material of many species collected by Hans Fruhstorfer, who mainly worked in Asia and South America (Baur and Coray 2004). Further, Yngve Sjöstedt, a Swedish biologist, who collected material during expeditions to Africa (Sjöstedt 1925), deposited many types in Hamburg. Carl Friedrich Brunner von Wattenwyl, one of the first specialists of the order Orthoptera and member of the Museum Godeffroy, worked with the collection of the ZMH (Weidner 1967). Hence, major parts of the Caelifera collection goes back to the Museum Godeffroy, a private collection belonging to a leader of a big trade company in the 18th century. Shortly after the Godeffroy Company bankrupted, most of the collections of the museum were sold and the zoological material was largely transferred to the ZMH in 1885 (Weidner 1967). Additional material now housed in the ZMH was collected during several expeditions by scientists of the Museum Hamburg (e.g. to Africa (Fischer 1884) and southwestern Australia (Michaelson 1908)) or was donated from researchers or private collectors. Many other important orthopterists worked with the collection and designated and deposited types; these include Boris Petrovitch Uvarov, D. Keith Mc. E. Kevan, Carl Stål, Grigory Yakovlevich Bey-Bienko, Willi Adolf Theodor Ramme, V. M. Dirsh, Jean-Guillaume Audinet-Serville, Ali Demirsoy, Anna Liana, Ignatio Bolivar, and Cornelis and Fer Willemse.

The collection currently houses 209 type specimens of Caelifera belonging to 128 species; 65 species are represented by holotypes (Table 1), three by lectotypes and one by a syntype. Besides a large number of paratypes and paralectotypes, the collection also houses topotypes of two species, which are not regulated by the International Code of Zoological Nomenclature, but are listed here for completeness. Type material of 120 species was recorded in previous catalogues presented by Weidner and Wagner (1968) and Weidner (1977). All previously recorded types were recovered, except the material of three species, which was already recorded as burned or missing by Weidner and Wagner (1968) (Cephalocoema teretiuscula, Brunner von Wattenwyl, 1890; Rhipipteryx striatipes, Chopard 1954; Tridactylus emarginatus, Chopard 1954). Further, the status of some species was not clear in Weidner’s catalogues and some species were recorded twice in the list; 115 of the species recorded by Weidner are also recorded here. Type material of 13 species has been deposited in the collection since the last catalogue.

Table 1.
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The table shows all type specimens currently housed in the collection of the ZMH including the original genus and species name, and the currently valid name (SYN = the species is now synonym of the species stated in this line). The type status is presented (H = holotype, P = paratype, PL = paralectotype (author as additional reference in last column), S = syntype, L = lectotype (author as additional reference in last column), T = topotype (recorded for completeness, not regulated by the taxonomic code). The number of present type specimens is given in the column ‘count’, followed by their gender (‘sex’). Author and publication year, as well as the collector, collection year and location are provided. The column ‘Weidner’, displays if the type had already been recorded in previous catalogues (Weidner & Wagner 1968; Weidner 1977; Y = yes, N = newly reported). Finally, we provide the reference (* Ref.) to the original description of each species; in case of lecto-, paralecto- or topotypes, author and year are presented (# Ref. number).

No. Genus Species Current taxonomy Type Count Sex Author, year Collector, year Location Weidner Ref.
Acrididae
1 Acanthacris ruficornis yemenita SYN Acanthacris ruficornis ruficornis H 1 1♂ Uvarov, 1936 C. Rathjens, 25.05.1931 Arabia, Yemen, Sanaa Y *27
2 Acrida curta SYN Acrida exaltata H 1 1♀ Uvarov, 1936 C. Rathjens, 1931 Arabia, Yemen, Sanaa Y *27
3 Adreppus tuberculatus - H 1 1♀ Sjöstedt, 1921 Museum Godeffroy Australia, Queensland, Peak-Downs Y *22
4 Apalacris gracilis - H 1 1♂ Willemse, 1936 H. Fruhstorfer, Nov. 1895 Sulawesi, Samanga, South Celebes Y *36
Apalacris gracilis - P 1 1♂ Willemse, 1936 H. Fruhstorfer, Nov. 1895 Sulawesi, Samanga, South Celebes Y *36
Apalacris gracilis - P 4 4♂ Willemse, 1936 H. Fruhstorfer, Jan. 1896 Sulawesi, Patunuang, South Celebes Y *36
5 Apalacris incompleta - H 1 1♀ Willemse, 1936 H. Fruhstorfer, Nov. 1895 Sulawesi, Samanga, South Celebes Y *36
6 Apoboleus affinis - H 1 1♂ Kevan, 1955 Hamb. Angola Expedition, 24.09.1952 Angola, Bengo Province, Piri (Dembos) Y *16
Apoboleus affinis - P 7 2♂ 5♀ Kevan, 1955 Hamb. Angola Expedition, 24.09.1952 Angola, Bengo Province, Piri (Dembos) Y *16
7 Apotropis affinis SYN Apotropis tricarinata H 1 1♀ Sjöstedt, 1921 Museum Godeffroy Australia, New South Wales, New Castle Y *22
Apotropis affinis SYN Apotropis tricarinata P 1 1♂ Sjöstedt, 1921 Museum Godeffroy Australia, New South Wales, New Castle Y *22
8 Asmara punctata Heteracris punctata H 1 1♂ Uvarov, 1936 C. Rathjens, Aug. 1931 Arabia, Yemen, Sanaa Y *27
Asmara punctata Heteracris punctata P 2 2♀ Uvarov, 1936 C. Rathjens, Aug. 1931 Arabia, Yemen, Sanaa Y *27
9 Asphingoderus uvarovites elazigi Asphingoderus elazigi P 2 1♂ 1♀ Demirsoy, 1979 A. Demirsoy, 15.07.1971 Turkey, Elazig N *7
10 Austracris guttulosa talawensis - P 2 1♂ 1♀ Sjöstedt, 1932 H. Fruhstorfer, 06.03.1898 Indonesia, Flores Y *24
11 Bibracte marginata - H 1 1♂ Willemse, 1931 Alb. Grubauer, 30.12.1902 Indonesia, Camp Jor, watershed between Perak and Pahang Y *32
12 Caparra amiculi SYN Urnisa rugosa H 1 1♀ Sjöstedt, 1921 Hamb. S. W. Austr. Expedition 1905 South-West Australia, north of Subiaco, Stat. 109 Y *22
13 Caperrala scabra - H 1 1♀ Sjöstedt, 1921 Hamb. S. W. Austr. Expedition, 07.09.1905 Australia, Edel Land Y *22
14 Cedarinia costata - P 1 1♀ Sjöstedt, 1921 Museum Stockholm, Mjöberg, 30.04.1921 Australia, Queensland, Tolga Y *22
15 Cedarinia scabra SYN Cedarinia vermiculata H 1 1♀ Sjöstedt, 1921 Museum Godeffroy Australia, Queensland, Rockhampton Y *22
16 Cedarinia variegata SYN Cedarinia vermiculata H 1 1♀ Sjöstedt, 1921 Museum Godeffroy Australia, Queensland, Peak-Downs Y *22
17 Clepsydria fuliginosa SYN Clepsydria historio P 4 2♂ 2♀ Sjöstedt, 1920 Museum Godeffroy, No. 80011 Australia, Queensland, Bowen Y *21
18 Desertaria cinnamomea SYN Desertaria cinnam P 1 1♀ Sjöstedt, 1920 Museum Stockholm, Mjöberg, 30.04.1921 Australia, Kimberley district Y *21
19 Dichroplus cliens - P 1 1♀ Stal, 1861 Museum Stockholm, 01.09.1922 Uruguay, Monte Video Y *26
20 Exarna includens - H 1 1♀ Walker, 1870 Australien, alte Sammlung ? Y *28
Exarna includens - P 1 1♂ Walker, 1870 Stal, No. 15844 Australia, Queens Land, Gayndah Y *28
21 Fenestra bohlsii - H 1 1♂ Giglio Tos, 1895 J. Bohls, 10.01.1895 Paraguay Y *10
22 Gastrimargus africanus orientalis SYN Gastrimargus africanus africanus P 1 1♀ Sjöstedt, 1928 Boltze, 1902 India, Hinterland of Assam Y *23
Gastrimargus africanus orientalis SYN Gastrimargus africanus africanus P 1 1♀ Sjöstedt, 1928 Boltze, 1902 East India Y *23
23 Gastrimargus africanus v. zebrata SYN Gastrimargus africanus africanus P 1 1♂ Sjöstedt, 1928 ? Kamerum, Sikumba near Maputa Y *23
Gastrimargus africanus v. zebrata SYN Gastrimargus africanus africanus P 1 1♀ Sjöstedt, 1928 ? Kamerum, Sikumba near Maputa Y *23
Gastrimargus africanus v. zebrata SYN Gastrimargus africanus africanus P 2 2♀ Sjöstedt, 1928 ? Kamerum, Mukonjjefarm am Mungofluß bei Mundame Y *23
Gastrimargus africanus v. zebrata SYN Gastrimargus africanus africanus P 1 1♀ Sjöstedt, 1928 ? Massai land Y *23
Gastrimargus africanus v. zebrata SYN Gastrimargus africanus africanus P 1 1♀ Sjöstedt, 1928 1893 Togo, Misahöhe Y *23
24 Gastrimargus brevipes v. abessina SYN Gastrimargus verticalis verticalis P 1 1♂ Sjöstedt, 1928 C. v. Erlangen, 7.12.1900 East Africa, Abessinien, Wonda (near Malke) Y *23
25 Gastrimargus clepsydrae SYN Gastrimargus crassicollis H 1 1♀ Sjöstedt, 1928 ? Transvaal, Johannesburg Y *23
26 Gastrimargus floresensis SYN Gastrimargus lombokensis H 1 1♂ Sjöstedt, 1928 H. Fruhstorfer, 05.06.1899 Indonesia, Flores Y *23
27 Gastrimargus musicus v. kimberleyensis SYN Gastrimargus musicus P 1 1♀ Sjöstedt, 1928 Museum Stockholm, Mjöberg, Jan. 1928 Australia, Kimberley District Y *23
28 Gastrimargus transvaalensis SYN Gastrimargus crassicollis P 2 2♀ Sjöstedt, 1928 Frau Will, 04.10.1902 South Africa, Johannesburg, Transvaal Y *23
29 Gastrimargus volkensi arabicus SYN Gastrimargus determinatus arabicus H 1 1♂ Uvarov, 1936 Dr. C. Rathjens, Aug. 1931 Arabia, Yemen, Sanaa Y *27
30 Goniaea acuta SYN Goniaea vocans P 1 1♂ Sjöstedt, 1921 Museum Godeffroy Australia, Queensland, Rockhampton Y *22
31 Goniaea glaucipes SYN Goniaea opomaloides H 1 1♀ Sjöstedt, 1921 H. Rolle, 26.10.1903 Australia, Victoria, Alexandra Y *22
32 Goniaea macronotum SYN Goniaea australasiae P 3 2♂ 1♀ Sjöstedt, 1921 ? Australia, Queensland, Peak-Downs Y *22
33 Goniaea vinaceipennis SYN Goniaea opomaloides H 1 1♀ Sjöstedt, 1921 Hamb. S. W. Austr. Expedition, 09-16.05.1905 Australia, Perth, north of Subiaco Y *22
34 Goniaeoidea jucunda SYN Goniaeoidea modesta H 1 1♂ Sjöstedt, 1921 Museum Godeffroy Australia, Queensland, Bowen Y *22
35 Heteropternis respondens insularis - H 1 1♂ Willemse, 1965 Le Moult, 1936 Java, Kangean Y *37
36 Hyalorrhipis urcmena grandis - P 2 1♂ 1♀ Bey-Bienko, 1960 Richter and Schäuffele, 11-21.04.1954 Iran, Belutschistan, Iranshar, 800m Y *2
37 Lobopteria pulchra - P 1 1♂ Sjöstedt, 1920 Museum Stockholm, Mjöberg, 30.04.1921 Australia, Queensland, Tolga Y *21
38 Loiteria rubripes - H 1 1♀ Sjöstedt, 1921 Museum Godeffroy, No. 15880 Australia, Queensland, Rockhampton Y *22
39 Opiptacris salomona - H 1 1♀ Ramme, 1941 Südsee-Exp. Wolf, 29.08.1909 Salomon Islands, Buka Y *20
40 Oxyblepta bohlsii Stenopola bohlsii H 1 1♀ Giglio-Tos, 1895 ? Paraguay Y *10
41 Oxytauchira elegans - H 1 1♂ Willemse, 1965 Le Moult, 1936 Java Y *37
42 Paratraulia elegantula SYN Tauchira rufotibialis P 1 1♂ Willemse, 1930 E. Jacobson, Feb. 1920 Sumatra, Westcoast, Tandjunggadang, 1200m Y *31
43 Pardillana ampla - P 1 1♂ Sjöstedt, 1920 Museum Stockholm, 30.04.1921 Australia, Kimberley District Y *21
44 Peakesia affinis SYN Desertaria lepida H 1 1♂ Sjöstedt, 1921 Museum Godeffroy Australia, Queens Land, Gayndah Y *22
45 Peakesia coeruleipes - P 2 1♂ 1♀ Sjöstedt, 1920 Museum Godeffroy Australia, Queensland, Peak-Downs Y *21
46 Perbellia brevialata - H 1 1♂ Sjöstedt, 1921 Museum Godeffroy Australia, Queensland, Bowen Y *22
47 Perbellia picta - P 1 1♂ Sjöstedt, 1920 Museum Stockholm, Mjöberg, 30.04.1921 Australia, Kimberley District Y *21
48 Phaulacridium nanum - P 1 1♀ Sjöstedt, 1921 Museum Godeffroy Australia, Sidney Y *22
49 Phlaeoba lombokensis Phlaeobella lombokensis P 2 2♂ Willemse, 1933 H. Fruhstorfer, May to June 1896 Indonesia, Lombok, Sapit, 2000m Y *34
50 Podismopsis relicta T 2 1♂ 1♀ Ramme, 1931 Willemse, 04.08.1965 Jugoslavia (=Montenegro), (Sbrija-Kosmet, Poc), Hajla-Cafe e Mustafes), 1900-2100m N #2 *19
51 Porraxia nana - H 1 1♀ Sjöstedt, 1921 Hamb. S. W. Austr. Exped. 12.05.1905 Australia, Tubiaco forest Y *22
52 Pseudogmo megalocephala SYN Pseudogmothela pallida H 1 1♂ Kevan, 1955 Hamb. Angola Expedition, Nr 2, 30.04.1953 Angola, Sanguengue, highlands of Nova Lisboa, 30km to Bela Vista Y *16
53 Pseudostauronotus brunneri Rhammatocerus brunneri H 1 1♂ Giglio Tos, 1895 ? Paraguay Y *10
54 Pycnodictya diluta - P 1 1♀ Ramme, 1929 Houy, Oct. 1912 Africa, Adamaua, near Garua and Rei Buba to Monti, 300-500m Y *18
55 Pycnodictya gracilis - P 4 2♂ 2♀ Uvarov, 1936 C. Rathjens, Aug. 1931 Arabia, Yemen, Sanaa Y *27
Pycnodictya gracilis - P 1 1♀ Uvarov, 1936 12.-14.03.1928 Arabia, Yemen, Sanaa-Manakha Y *27
Pycnodictya gracilis - P 2 1♂ 1♀ Uvarov, 1936 C. Rathjens, Apr. 1931 Arabia, Yemen, Hadja Y *27
56 Rapsilla fusca - P 1 1♂ Sjöstedt, 1921 Museum Godeffroy, No. 2719 Australia, New-South-Wales Y *22
Rapsilla fusca - P 1 1♀ Sjöstedt, 1921 Nelles, 24.05.1898 Australia, New-South-Wales Y *22
57 Rapsilla testacea SYN Rapsilla fusca H 1 1♀ Sjöstedt, 1921 Nelles, 24.05.1898 Australia, New-South-Wales Y *22
58 Rhitzala modesta - P 1 1♀ Sjöstedt, 1921 Museum Godeffroy, No. 15893 Australia, Sidney Y *22
59 Schistocerca centralis - P 3 2♂ 1♀ Dirsh, 1974 R. Paessler, 04-06.05.1912 Costa Rica, Punta Arenas Y *9
Schistocerca centralis - P 3 2♂ 1♀ Dirsh, 1974 R. Paessler, 04-06.05.1912 Central America, Corinto, 12°29’N 84°14‘25W Y *9
60 Schizobothrus flavovittatus - P 1 1♀ Sjöstedt, 1921 Hamb. S. W. Austr. Expedition 1905 Australia, Perth, north of Subiaco, Stat. 109 Y *22
61 Sphodromerus rathjensi rathjensi - H 1 1♀ Uvarov, 1936 C. Rathjens, Sep. 1931 Arabia, Yemen, Sanaa Y *27
62 Stenocatantops cornelii - P 4 2♂2♀ Willemse, 1968 Le Moult, 1936 Java, Semarang Y *38
Stenocatantops cornelii - P 2 1♂1♀ Willemse, 1968 Le Moult, May 1936 Java, Mt. Djampang Y *38
Stenocatantops cornelii - P 1 1♀ Willemse, 1968 Le Moult, May 1936 Java, Radjamandala Y *38
Stenocatantops cornelii - P 1 1♀ Willemse, 1968 Le Moult, May 1936 Java, Penandjoeng Y *38
63 Stropis nigrovitellina - P 1 1♂ Sjöstedt, 1920 Museum Stockholm, Mjöberg, 30.04.1921 Australia, Kimberley District Y *21
64 Stropis viridis SYN Stropis nigrovitellina P 1 1♀ Sjöstedt, 1920 Museum Stockholm, Mjöberg, 30.04.1921 Australia, Kimberley District Y *21
65 Terpillaria picta SYN Macrocara conglobata P 1 1♀ Sjöstedt, 1921 Museum Godeffroy, No. 7989 Australia, Queensland, Bowen Y *22
66 Terpillaria pulchra - P 1 1♂ Sjöstedt, 1921 Museum Stockholm, Mjöberg, 30.04.1921 Australia, Kimberley District Y *21
67 Trachypetrella kosswigiana Trachypetrella anderssonii H 1 1♀ Weidner, 1964 H. Rolle, 14.06.1905 Africa, Tanganyika, Lake Tanganyika Y *29
68 Urnisa gutulosa minor L 1 1♀ Sjöstedt, 1921 W. Wölting, 1907 West Australia, Fremantle Y #1 *22
Urnisa gutulosa minor PL 1 1♀ Sjöstedt, 1921 W. Wölting, 1907 West Australia, Fremantle Y #1 *22
69 Valanga irregularis carolinensis Valanga nigricornis carolinensis H 1 1♀ Sjöstedt, 1932 Museum Godeffroy Caroline Islands, Ruck Y *24
70 Valanga irregularis v. signata - P 2 1♂ 1♀ Sjöstedt, 1921 Museum Godeffroy Australia, Queensland, Rockhampton Y *22
71 Valanga modesta - P 1 1♀ Sjöstedt, 1921 Museum Godeffroy Australia, Queens Land, Gayndah Y *22
72 Valanga nigricornis v. aroensis - P 1 1♂ Sjöstedt, 1932 Brunner von Wattenwyl, 1889 Indonesia, Aru-Islands Y *24
73 Valanga nobilis miokoana - H 1 1♀ Sjöstedt, 1932 ? Mioko Y *24
74 Valanga rubrispinarum - P 1 1♀ Sjöstedt, 1936 Museum Godeffroy Australia, Queensland, Rockhampton Y *25
75 Valanga soror - L 1 1♂ Sjöstedt, 1936 Museum Goddefroy Australia, Queens Land, Gayndah Y *25 #1
Valanga soror - P 1 1♀ Sjöstedt, 1936 Museum Goddefroy Australia, Queens Land, Gayndah Y *25
76 Valanga viridipes SYN Valanga nigricornis carolinensis H 1 1♀ Sjöstedt, 1932 ? Caroline Islands, Ruck Y *24
77 Xenocatantops dirshi - P 6 6♀ Willemse, 1968 Le Moult, 1936 Java, Kangean Y *38
78 Xenocatantops humilis T 1 1♂ Serville, 1838 Le Moult, 1936 Java, Penandjoeng N #2 *1
Xenocatantops humilis - T 2 2♂ Serville, 1838 Le Moult, 1936 Java, Mt. Djampang N #2 *1
Xenocatantops humilis - T 2 2♀ Serville, 1838 Le Moult, Oct. 1936 Java, Radjamendal N #2 *1
Xenocatantops humilis - T 2 2♀ Serville, 1838 Le Moult, Oct. 1936 Java N #2 *1
79 Zebratula flavonigra - P 1 1♀ Sjöstedt, 1920 Museum Stockholm, Mjöberg, 30.04.1921 Australia, Kimberley District Y *21
Chorotypidae
80 Eupatrides signatus - H 1 1♂ Willemse, 1935 H. Fruhstorfer, Feb. 1896 Sulawesi, Samanga, South Celebes, Bua-Kraeng, 5000m Y *35
Morabidae
81 Callitala major - L 1 1♂ Sjöstedt, 1936 Museum Godeffroy Australia, Queensland, Rockhampton Y #1 *22
Callitala major - PL 1 1♀ Sjöstedt, 1936 Museum Godeffroy Australia, Queensland, Rockhampton Y #1 *22
82 Moraba amiculi Heide amiculi P 2 2♀ Sjöstedt, 1921 Museum Godeffroy Australia, New-South-Wales, Sidney Y *22
83 Moraba laticornis Swanea laticornis H 1 1♀ Sjöstedt, 1921 Hamb. S. W. Austr. Expedition 1905 Australia, Bunswick, Stat. 139 Y *22
84 Moraba obscura - P 1 1♀ Sjöstedt, 1921 Hamb. S. W. Austr. Expedition 1905 Australia, Lion Mill, Stat. 139 Y *22
85 Moraba parva Amangu parva H 1 1♂ Sjöstedt, 1921 Hamb. S. W. Austr. Expedition 1905 Australia, Perth, Karrakatta, Stat. 139 Y *22
Ommexechidae
86 Aucacris hebardi SYN Aucacris eumera H 1 1♀ Willemse, 1931 O. Voelckler, 06.02.1899 Chile Y *33
Pamphagidae
87 Paranocaracris citripes bicoloripes f. marani SYN Nocarodes bicoloripes P 1 1♂ Demirsoy, 1973 Zoologisch Botanische Exkursion 1964 Anatolia, Erciyas Dag Y *6
88 Paranocarodes fieberi anatoliensis Paranocarodes anatoliensis P 1 1♀ Demirsoy, 1973 A. Demirsoy, 17.07.1949 Turkey, Antalya, Tahtali Dagi, ca. 1500m Y *6
89 Paranothrotes kosswigi SYN Nocaracris P 1 1♀ Demirsoy, 1973 D. D. Hasan , 20.06.1968 Turkey, Erzincan-Kamaliye-Yuvaköyü (Bügelek Kayasi) Y *6
Proscopiidae
90 Altograciosa mirabilis - H 1 1♂ Liana, 1980 J. Hubrich Argentina, Province Cordoba, La Granja, near Alta Gracia N *17
91 Anchotatus peruvianus - S 1 1♀ Brunner von Wattenwyl, 1890 Brunner von Wattenwyl, 1889 Peru Y *5
92 Anchotatus similis - H 1 1♂ Liana, 1980 F. Ohaus, 20.09.1905 Ecuador, Zaruma N *17
Anchotatus similis - P 1 1♀ Liana, 1980 F. Ohaus, 20.09.1905 Ecuador, Zaruma N *17
93 Apioscelis gracilis SYN Apioscelis bulbosa P 2 1♂1♀ Brunner von Wattenwyl, 1890 11.12.1889 Peru, Alto Amazonas Province Y *5
94 Astroma carbonelli - H 1 1♂ Liana, 1980 R. Paessler, 17.03.1914 Bolivia, Oruro N *17
95 Astroma plicatum - H 1 1♀ Liana, 1980 Meteor Expedition, Dec. 1925 Cordillera, Poterillos near Mendoza N *17
Astroma plicatum - P 1 1♀ Liana, 1980 Meteor Expedition, Dec. 1925 Cordillera, Poterillos near Mendoza N *17
96 Callangania brunneri - H 1 1♂ Liana, 1980 Dr. O. Staudinger, 20.05.1899 Peru, Callanga N *17
97 Cephaloscopia grisea Anchotatus griseus H 1 1♀ Liana, 1973 R. Paessler, 02-08.10.1914 Chile, Coronel N *17
Cephaloscopia grisea Anchotatus griseus P 1 1♀ Liana, 1973 Jacobsen?, 12.05.1899 Chile, Concepción N *17
98 Cephaloscopia weidneri Anchotatus weidneri H 1 1♀ Liana, 1973 Sammelreise 1936, 21.04.1936 South Peru, Cayara, ca. 3200m N *17
99 Corynorhynchus subtilis - H 1 1♂ Liana, 1980 F. Ohaus Brasilia, Petropolis N *17
Corynorhynchus subtilis - P 2 1♂1♀ Liana, 1980 F. Ohaus Brasilia, Petropolis N *17
100 Corynorhynchus tibarisae - H 1 1♂ Liana, 1980 F. Ohaus, 20.10.1926 Costa Rica S Paulo, Tibirisá N *17
Corynorhynchus tibarisae - P 2 1♂1♀ Liana, 1980 F. Ohaus, 20.10.1926 Costa Rica S. Paulo, Tibirisá, near Rio Parana N *17
Corynorhynchus tibarisae - P 1 1♀ Liana, 1980 F. Ohaus, 03.12.1926 Brasil, Rio de Janeiro, 900-1000m N *17
101 Prosarthria teretrirostris - P 1 1♀ Brunner von Wattenwyl, 1890 H. Rode Venezuela Y *5
102 Pseudoanchotatus jaczewski Pseudoanchotatus jacsewskii H 1 1♀ Liana, 1980 C. Bock, 23.07.1903 Bolivia, Rio Sango N *17
103 Tetanorhynchus longirostris - P 1 1♀ Brunner von Wattenwyl, 1890 H. Fruhstorfer, 12.02.1889 Brazilia, St. Catharina Province Theresopolis Y *5
104 Tetanorhynchus propinquus - P 1 1♀ Brunner von Wattenwyl, 1890 Killmann?, 07.11.1889 Brazilia, St. Catharina Province, Joinville Y *5
105 Xenium saburrae - H 1 1♀ Liana, 1980 ? Chile N *17
Pyrgomorphidae
106 Brunniella antistes - H 1 1♀ Bolivar, 1905 Alte Sammlung Phillippines, Patria? Y *4
107 Dictyophorus grisea f. brunni - H 1 1♀ Bolivar, 1904 Hamb. Geographische Gesellschaft, Fischer, 1884 Africa, Massai-Land Y *3
108 Dictyophorus spumans v. pulchra - H 1 1♂ Bolivar, 1904 E. Heyne, 23.05.1900 South Africa, Natal, Marianhill Y *3
109 Monistria cordata SYN Greyacris profundesulcata H 1 1♀ Sjöstedt, 1921 Museum Godeffroy Australia, Queensland, Bowen Y *22
110 Monistria latevittata - H 1 1♀ Sjöstedt, 1921 Hamb. S. W. Austr. Expedition, 23-25.09.1905 South-West-Australia, Serpentine Y *22
111 Monistria ligata - H 1 1♀ Bolivar, 1904 ? Australia, New-South-Wales, Sidney Y *3
112 Monistria maculicornis - P 1 1♀ Sjöstedt, 1921 Hamb. S. W. Austr. Expedition, 23-25.09.1905 W Austr., Serpentine Y *22
113 Monistria mastax - H 1 1♂ Sjöstedt, 1921 Museum Godeffroy Australia, Queensland, Rockhampton Y *22
114 Paraspena yemenita H 1 1♂ Uvarov, 1936 C. Rathjens, Aug. 1931 Arabia, Yemen, Sanaa Y *27
Paraspena yemenita P 1 1♀ Uvarov, 1936 C. Rathjens, Aug. 1931 Arabia, Yemen, Sanaa Y *27
115 Pyrgomorpha minuta - P 1 1♀ Kevan, 1963 E. R. Helwig, 03.01.1939 Africa, Cape Province, Cape Flats, False Bay, Cape Town Y *8
116 Tanita longiceps Protanita longiceps H 1 1♀ Bolivar, 1904 Stuhlmann, 12.09.1888 Africa, Tanganyika, Kihenga Y *3
Tetrigidae
117 Allotettix otumboides - P 2 2♂ Günther, 1939 O. Staudinger, 20.05.1899 Peru, Callanga Y *13
118 Bolivaritettix difficilis - P 2 2♀ Günther, 1939 H. Fruhstorfer, 10.02.1894 Java Y *13
119 Chiriquia compressa - P 1 1♂ Günther, 1939 R. Haensch, 30.04.1903 Ecuador, Santa Inez Y *13
120 Hyboella overbecki - P 1 1♀ Günther, 1939 Museum Marburg, 24.07.1901 Java Y *13
121 Hyboella perakensis - H 1 1♀ Günther, 1939 A. Grubauer, 30.12.1902 Indonesia, Camp Jor, watershed between Perak and Pahang Y *13
122 Loxilobus brunneri - H 1 1♀ Günther, 1938 H. Schrader, 1913 Nieder-Burma, near Rangoon a. Unterl. D. Irwati Y *12
123 Loxilobus kraepelini - H 1 1♀ Günther, 1938 K. Kraepelin, 24.02-12.03.1904 Java, Buitenzorg Y *12
124 Paratettix amplus - P 1 1♂ Sjöstedt, 1921 Museum Goddefroy, No. 15853 Australia, Queensland, Rockhampton Y *22
125 Paratettix meridionalis weidneri SYN Paratettix meridionalis P 1 1♂ 2♀ Karaman, 1965 T. Becker, 1901 Canary Islands, Tenerife Y *15
126 Systolederus carli celebensis SYN Systolederus ophthalmicus P 2 2♂ Günther, 1937 H. Fruhstorfer, Nov. 1895 Sulawesi, Samanga, South Celebes Y *11
Tridactylidae
127 Afrotridactylus meridianus - P 3 2♂1♀ Günther, 1994 W. Krieg, 1933-1934 South West Afrika, Okundua near Okahandja N *14
Trigonopterygidae
128 Systella borneensis - P 1 1♀ Willemse, 1930 H. Rolle, 25.11.1904 Borneo, Kinabalu, 1500m, Watersadt Y *30
129 Systella dusmeti - PL 1 1♂ Bolivar, 1905 H. Rolle, 25.11.1904 Borneo, Kinabalu, 1500m, Watersadt Y *4
Systella dusmeti - PL 1 1♂ Bolivar, 1905 H. Fruhstorfer, 05.06.1899 North Borneo Y *4
Systella dusmeti - PL 1 1♀ Bolivar, 1905 A. Speyer, 20.11.1899 North Borneo Y *4
130 Systella philippensis Systella siccifolia H 1 1♂ Walker, 1870 Patria, Alte Sammlung Patria? Y *28

Literature: #1 Key (1958); #2 Willemse (1964); *1 Audinet-Serville (1839); *2 Bey-Bienko (1960); *3 Bolívar (1904); *4 Bolívar (1905); *5 Wattenwyl (1890); *6 Demirsoy (1973); *7 Demirsoy (1979); *8 Dirsh (1963); *9 Dirsh (1974); *10 Giglio-Tos (1895); *11 Günther (1937); *12 Günther (1938); *13 Günther (1939); *14 Günther (1994); *15 Karaman (1965); *16 Kevan (1955); *17 Liana (1980); *18 Ramme (1929); *19 Ramme (1931); *20 Ramme (1941); *21 Sjöstedt (1920); *22 Sjöstedt (1921); *23 Sjöstedt (1928); *24 Sjöstedt (1932); *25 Sjöstedt (1936); *26 Stål (1851); *27 Uvarov (1936); *28 Walker (1870); *29 Weidner (1964); *30 Willemse (1930)a; *31 Willemse (1930)b; *32 Willemse (1931)a; *33 Willemse (1931)b; *34 Willemse (1933); *35 Willemse (1935); *36 Willemse (1936); *37 Willemse (1965); *38 Willemse (1968)

In addition to ZMH types reported here, other specimens originally designated as type, but not formally published, are present in the collection (Fig. 1d); this material is not listed here. An example comes from the genus Paratettix: Günther (1979) mentioned material of the genus, which likely belongs to an undescribed species. This material is marked as such in the collection, but a new species was never formally described. Further, it is likely that additional types, not marked as such, are present; especially the material of the Collection Godeffroy is likely to contain additional type specimens. Experts of the respective groups are needed to investigate and work with the collection. This paper is one of the first in a row of reports to be published in the next years to present updated type catalogues of the insect collection of the ZMH (now part of the CeNak).

Figure 1.

Shown are a) a cotype of Gastrimargus musicus v. kimberleyensis Sjöstedt, 1928; b) the holotype of Adreppus tuberculatus Sjöstedt, 1921; c) the holotype of Callangania brunneri Liana, 1980; and d) the tentative type of a Tetrigid, identified as unknown, but not formally described by Kurt Günther and their labels.

Acknowledgement

We want to thank Holger Braun, Bruno Massa and an anonymous reviewer for valuable comments on a previous version of this manuscript. This project was funded by the Orthopterists’ Society and by the Gesellschaft der Freunde und Förderer des Zoologischen Museums Hamburg.

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