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An illustrated catalogue of the type specimens of Lepidoptera (Insecta) housed in the Zoological Museum Hamburg (ZMH): Part I. superfamilies Hepialoidea, Cossoidea, and Zygaenoidea
expand article infoReza Zahiri§, Gerhard Tarmann|, Konstantin A. Efetov, Hossein Rajaei#, Maryam Fatahi, Matthias Seidel, Birgit Jaenicke¤, Thure Dalsgaard, Marcy Sikora, Martin Husemann
‡ University of Hamburg, Hamburg, Germany
§ Canadian Food Inspection Agency, Ottawa, Canada
| Naturwissenschaftliche Sammlungen, Sammlungs- und Forschungszentrum der Tiroler Landesmuseen, Hall, Austria
¶ V. I. Vernadsky Crimean Federal University, Simferopol, Russia
# State Museum of Natural History Stuttgart, Stuttgart, Germany
¤ Museum für Naturkunde, Berlin, Germany
Open Access

Abstract

Zoological collections remain the main archive of animal biodiversity on Earth, and especially in times of large-scale declines of many groups represent important resources for biodiversity and conservation research. The most important individuals of these collections are the type specimens, which represent the original and unique reference for a species. While a full digitization of most museum collections currently remains utopic, lists of types are an essential resource for researchers to perform taxonomic revisions. Here, we provide an updated type catalogue of the lepidopteran superfamilies Hepialoidea, Cossoidea and Zygaenoidea deposited in the Zoological Museum of Hamburg (ZMH). In this paper, we report 259 specimens labelled as “types” belonging to 36 infrasubspecific taxa (invalid under the ICZN codes), 34 subspecies (16 of valid status) and 13 species (all of valid status). We present a full bibliography of the original descriptions and illustrations for all taxa, aiming to provide a comprehensive taxonomic guide to this collection.

Key Words

Biodiversity, CeNak, Herbert Weidner, ZMH, digitization, Georg Warnecke, Hanan Bytinski-Salz

Introduction

Natural history collections are the main archives of biodiversity. They focus on collecting, maintaining and documenting natural specimens, a crucial task in times of biodiversity decline. The Zoological Museum of Hamburg (ZMH) holds large collections across all animal groups. With approximately five million specimens and several thousand primary types, the entomological collection is among the most important natural history archives in Germany.

The origins of the current collection of the Zoological Museum Hamburg (ZMH) can be traced back to the first half of the 19th century, as a result from the initiative and interest in natural history of local civilians, merchandisers, traders, and owners of seagoing vessels (Glaubrecht 2018). However, much of the collections of holometabolous insects were destroyed by the incendiary bombing of Hamburg in 1943 (Weidner 1979). Thus, as all documents such as entry books, counting lists and card catalogs were lost and the former magnitude of the collection remains difficult to describe. However, much of the earlier new entries after WW2 are described in Weidner (1974), with the largest and most important collections which have been obtained being those of Georg Warnecke (1943 and 1963), Richard Jänig (1949), Fritz Diehl (1950–1972), Hermann Rödinger (1952), the Jungius Gesellschaft (1956), Hanan Bytinski-Salz (1960), Eduard Feldtmann (1960), Günter Albers (1972), the Altona Museum (1965, 1973, 1976), and Mathilde Linz (1974). More recently the largest newly obtained collections were those of Hans-Jürgen Kelm (2003), Walter Baltruweit (2013), Jörg Roloff (2014), Thomas Tischler (2017), Bernd Heinze (2020), and Karl-Heinz Müller-Köllges (2020). Besides these large collections, many medium and small collections were obtained by the museum. Hence, despite the considerable loss in WW2, presently the ZMH hosts ca. 5 million insect specimens, being placed among the five largest entomological collections in Germany. About 1 million specimens represent the Lepidoptera. However, despite their importance, the entomological collections in Hamburg are not well documented, nor digitized.

Herbert Weidner, a former curator of the collection had published an important series of papers from the 60s until the late 70s, where he documented the collections in much detail (first catalogue – Weidner 1962, last catalogue – Weidner 1979, and the Lepidoptera catalogue Weidner 1974). However, since the work of Weidner more than 40 years have passed without further documentation. Further, Weidner’s catalogues were all published in German, hence, are difficult to access for international researchers and lack family group classification, making it difficult to work with them. Therefore, a comprehensive effort to revise the type catalogues has been initiated to give an update on the type material housed in the ZMH in English language (Sartori et al. 2016; Dey and Husemann 2018a, b, Harms and Dupérré 2018; Monod et al. 2019; Henningsen et al. 2020; Simoes et al. in press).

To make the Lepidoptera collection more accessible to researchers, we performed a detailed inventory initiated by Hossein Rajaei in 2013. As a part of this inventory, the whole collection was thoroughly searched and types were moved to a separate, specially protected type collection. All type specimens were compared with the original descriptions, photographed, databased and relabeled, if required. The original high-quality photographs are available online through the FUNDus! Portal of the University of Hamburg (https://www.fundus.uni-hamburg.de/). The complete data of the type material is provided as a table (Suppl. material 1: Table S1) in this catalogue.

Material and methods

Photography. All images were taken with a digital imaging system with stacking capabilities. The images where either taken with a Canon EOS 5DSR with a 65 mm or 100 mm lens and stacked with Zerene Stacker (PMax algorithm) or with a Canon EOS 6D with a 55 mm or 65 mm lens and stacked with Helicon Focus 5.3.

Taxonomy. All taxonomic changes were made after studying the relevant original description of the type material. Synonyms that have already been proposed are listed and the type material was examined. Classification used herein follows van Nieukerken et al. (2011). The taxonomy of the family Zygaenidae is following the last revisions of the family (Hofmann and Tremewan 1996; Efetov 2001; Efetov and Tarmann 1995, 2017). For a full list of synonyms, please refer to the above-mentioned catalogues.

Type and original specimen material. Type and original specimen localities are cited in their original spelling or with additional remarks (in square brackets), if required. All data are transcribed verbatim as they appeared on the respective labels. For unreadable data aquestion mark (?) is used. Double slash (//) separates data on different labels, a single slash (/) separates lines within each label, and semi-colon (;) separates different specimens. Additional comments are placed in square brackets.

The ICZN (1999) terminologies (e.g., holotype, syntype, lectotype, neotype, paratype, paralectotype, allotype, cotype and etc.) are available at the ICZN website (http://iczn.org/content/Glossary) and articles cited in the text can be found below:

Infrasubspecific taxon – a taxon at lower rank than that of subspecies. The names of such taxa are not regulated by the Code.

Article 45.6.1. – A name is infrasubspecific if its author expressly gave it infrasubspecific rank, or if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity;

Article 45.6.2. – A name is deemed to be infrasubspecific if its author used one of the terms “aberration”, “ab.” or “morph”.

Article 45.6.3. – A name is deemed to be infrasubspecific if it was first published after 1960 and the author expressly used one of the terms “variety” or “form” (including use of the terms “var.”, “forma”, “v.” and “f.”);

Article 45.6.4. – A name is subspecific if first published before 1961 and its author expressly used one of the terms “variety” or “form” (including use of the terms “var.”, “forma”, “v.” and “f.”), unless its author also expressly gave it infrasubspecific rank, or the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, in which case it is infrasubspecific;

Article 72.7. – Name-bearing types of nominal species-group taxa denoted by new replacement names (nomina nova). If an author proposes a new species-group name expressly as a replacement (a nomen novum) for an earlier available one, then the two names are objective synonyms; both the nominal taxa they denote have the same name-bearing type despite any simultaneous restriction or application of the new replacement name (nomen novum) to particular specimens or any contrary designation of type, or any different taxonomic usage of the new replacement name.

Type specimens deposited at the Zoological Museum Hamburg (ZMH)

Superfamily Hepialoidea Stephens, 1829

Hepialoidea (the ghost moths and their relatives), with 629 described species worldwide, is the earliest diverging lepidopteran clade to exhibit any significant degree of species diversity today (Nielsen et al. 2000). The family Hepialidae is the most diverse group among other five families within the superfamily and comprises 606 species in 62 genera. In contrast to the remainder of the pre-Ditrysian Lepidoptera, many of the species are large and spectacular moths (Nielsen et al. 2000).

Family Hepialidae Stephens, 1829

Phymatopus hecta ab. inversa (Bytinski-Salz, 1939)

Hepialus hecta ab. inversa Bytinski-Salz, 1939: Ent. Rec. 51: 85.

Original material examined

Labelled as “Holotype” (ZMH 61345), 1♀ “St. Amata / Lettland / 16.7.1933 / leg. W. Brandt // Hepilus / hecta ab. / inversa / ByS / Holotypus f // ab. inversa / ByS // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61345”.

Original locality

St. Amata, Lettonia [Latvia].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1939) proposed this name as an aberration of a H. hecta, therefore it is deemed to be infrasubspecific, which is hence unavailable (Article 45.6.2, ICZN 1999). The original locality is Latvia, not Britain as mentioned in Nielsen et al. (2000).

Phymatopus hecta ab. confluens (Bytinski-Salz, 1939)

Hepialus hecta ab. confluens Bytinski-Salz, 1939: Ent. Rec. 51: 84.

Original material examined

Labelled as “Holotype” (ZMH 61346), 1♂. Thundersley (Essex, Britain), 06.vii.1911, leg. A.T. Stiff. (Fig. 2). “Hepialus / hecta / ab. confluens / ByS / Holotypus m // A.T. Stiff, / Thundersley, SX. / July 6. 1911. // ab. confluens / ByS. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61346”.

Original locality

Thundersley [Essex, Britain].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1939) proposed this name as an aberration of a H. hecta, therefore it is deemed to be infrasubspecific, which is hence unavailable (Article 45.6.2, ICZN 1999).

Pharmacis fusconebulosus ab. latefasciatus (Bytinski-Salz, 1939)

Hepialus fusconebulosus ab. latefasciatus Bytinski-Salz, 1939: Ent. Rec. 51: 83.

Original material examined

Labelled as “Holotype” (ZMH 61347), 1♀ (Fig. 3). “Hepialius / fuscunebulosus / ab. latefasciatus / ByS / Holotypus f // ab. latefasciatus / ByS. // Pitcaple. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61347”.

Original locality

Pitcaple [Scotland].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1939) proposed this name as an aberration of a H. hecta, therefore it is deemed to be infrasubspecific, which is hence unavailable (Article 45.6.2, ICZN 1999).

Pharmacis fusconebulosus ab. ornatus (Bytinski-Salz, 1939)

Hepialus fusconebulosus ab. ornatus Bytinski-Salz, 1939: Ent. Rec. 51: 83.

Original material examined

Labelled as “Holotype” (ZMH 61348), 1♂, leg. A.J. Hodges (Fig. 4). “ab. ornatus / ByS // A.J.Hodges / Stiga- // Hepialius / fuscunebul. / ab. ornatus ByS // Holotypus m // ZMH 61348”.

Original locality

Sligo [Ireland].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1939) proposed this name as an aberration of a H. fusconebulosus, therefore it is deemed to be infrasubspecific, which is hence unavailable (Article 45.6.2, ICZN 1999).

Hepialus humuli ab. roseoornata (Bytinski-Salz, 1939)

Hepialus humuli ab. roseoornata Bytinski-Salz, 1939: Ent. Rec. 51: 81.

Original material examined

Labelled as “Holotype” (ZMH 61349), 1♀, xii.1914, leg. A.T. Stiff (Fig. 5). “Tavistock / A.T. Stiff / July 1914 // ab. roseoornata / ByS. // Hepialus / humuli ab. / roseoornata / ByS / Holotypus f // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61349”.

Original locality

Tavistock [Britain].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1939) proposed this name as an aberration of H. humuli, therefore it is deemed to be infrasubspecific, which is hence unavailable (Article 45.6.2, ICZN 1999).

Hepialus humuli thulensis f. uniformis (Bytinski-Salz, 1939)

Hepialus humuli thulensis f. uniformis Bytinski-Salz, 1939: Ent. Rec. 51: 82.

Original material examined

Labelled as “Cotype” (ZMH 61358 – 61359), 2♂♂, Shetland Isl., leg. 1909 (Fig. 7). Shetlands / 09 // ab. uniformis / Bys // Hepialus / humuli / ab. uniformis / Bys / CoTypus m // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61358”; “Shetlands / 09 // Hepialus / humuli / ab. uniformis / Bys / Cotypus m // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61359”.

Original locality

Shetland [Britain].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1939) proposed this name as a form of a subspecies. According to Article 45.6.4., the author expressly gave it infrasubspecific rank, in which case it is infrasubspecific, and hence unavailable.

Hepialus humuli thulensis f. albida (Bytinski-Salz, 1939)

Hepialus humuli thulensis f. albida Bytinski-Salz, 1939: Ent. Rec. 51: 82.

Original material examined

Labelled as “Cotype” (ZMH 61350 – 61356), 7♂♂, leg. A.J. Hodges, vii.1899, Shetland, 1909 (Fig. 6). “humuli – / hetlandica / 5 Shetland // Cotypus thulenis / t. albisa Bys. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61350”; “Shetlands / 09 // Cotypus / thulensis / f. albida ByS. // Coll. Bytinski-Salz / Eing. Nr. 20 – 60 // ZMH 61351”; “Shetlands / 09 // Cotypus / thulensis / f. albida ByS. // ZMH 61352”; “Shetlands / 09 // Cotypus / thulensis / f. albida ByS. // Coll. Bytinski-Salz / Eing. Nr. 20–60 // ZMH 61353”; “A.J.Hodges / N.UNST / Y.99. N.B // Cotypus / thulensis / f. albida ByS. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61354”; “A.J.Hodges / Y. 99 Unst / Shitlands // Cotypus / thulensis / f. albida Bys. // Coll. Bytinski-Salz / Eing. Nr. 20–60 // ZMH 61355”; “ A.J.Hodges / N.UNST / Y.99. N.B // f. albida / Bys. // Cotypus / thulensis / f. albida ByS. // Coll. Bytinski-Salz / Eing. Nr. 20–60 // ZMH 61356”.

Original locality

Shetland [Britain].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1939) proposed this name as a form of a subspecies. According to Article 45.6.4., the author expressly gave it infrasubspecific rank, in which case it is infrasubspecific, and hence unavailable.

Phymatopus hecta ab. ornata (Bytinski-Salz, 1939)

Hepialus hecta ab. ornata Bytinski-Salz, 1939: Ent. Rec. 51: 85.

Original material examined

Labelled as “Holotype” (ZMH 61357), 1♂ (Fig. 8). Thundersley, SX (Essex, Britain), 04.vii.1911, leg. A.T. Stiff. “Hepialus / hecta / ab. ornata / ByS / Holotypus m // A.T. Stiff, / Thundersley, SX. / July 4 1911. // Coll. Bytinski-Salz / Eing. Nr. 20–60 // ab. ornata / ByS. // ZMH 61357”.

Original locality

Thundersley [Essex, Britain].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1939) proposed this name as an aberration of a H. hecta, therefore it is deemed to be infrasubspecific, which is hence unavailable (Article 45.6.2, ICZN 1999).

Korscheltellus lupulina ab. latemarginatus (Bytinski-Salz, 1939)

Hepialus lupulinus ab. latemarginatus Bytinski-Salz, 1939: Ent. Rec. 51: 84.

Original material examined

Labelled as “Cotype” (ZMH 61360−60361), 2♂♂. London Dist. And Wicken, leg. A.J. Hodges (Fig. 9). “London Dist. // ab. latemarginatus / ByS // ZMH 61360 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // Hepialus / lupulina / ab. latemar- / ginatus ByS / Cotypus m”; “A.J.Hodges / Wicken // Hepialus / lupulina / ab. latemar- / ginatus / ByS / Cotypus m // Coll. Bytinski-Salz / Eing. Nr. 20–60 // ZMH 61361”.

Figures 1–9.

1. Phymatopus hecta ab. inversa (Bytinski-Salz, 1939); 2. Phymatopus hecta ab. confluens (Bytinski-Salz, 1939); 3. Pharmacis fusconebulosus ab. latefasciatus (Bytinski-Salz, 1939); 4. Pharmacis fusconebulosus ab. ornatus (Bytinski-Salz, 1939); 5. Hepialus humuli ab. roseoornata (Bytinski-Salz, 1939); 6. Hepialus humuli thulensis f. uniformis (Bytinski-Salz, 1939); 7. Hepialus humuli thulensis f. albida (Bytinski-Salz, 1939); 8. Phymatopus hecta ab. ornata (Bytinski-Salz, 1939); 9. Korscheltellus lupulina ab. latemarginatus (Bytinski-Salz, 1939).

Original localities

London district and Wicken [Britain].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1939) proposed this name as an aberration of a H. lupulinus, therefore it is deemed to be infrasubspecific, which is hence unavailable (Article 45.6.2, ICZN 1999).

Superfamily Cossoidea Leach, 1815

The superfamily Cossoidea currently contains seven families (i.e., Brachodidae, Cossidae, Dudgeoneidae, Metarbelidae, Ratardidae, Castniidae, and Sesiidae) (van Nieukerken et al. 2011) with type representatives from the families Cossidae, Brachodidae, Metarbelidae, and Sesiidae in ZMH collection. They are generally concealed feeders and have spiny pupae with moveable segments to allow them to extrude out of their exit holes in stems and trunks during emergence of the adult (Edwards et al. 1999). The clearwing moths (Sesiidae) represent the largest radiation within the superfamily with about 1,400 valid species distributed in 154 genera (van Nieukerken et al. 2011). The carpenter moths (Cossidae) is the second largest group with about 1000 described species in 151 genera (Yakovlev 2011). The family Metarbelidae with more than 240 described species in 32 genera have their highest diversity in the Afrotropics (Lehmann 2019; Yakovlev and Zolotuhin 2020). However, the family is among the least known of Lepidoptera due their extreme rarity and rather cryptic appearance (Lehmann 2019).

Family Cossidae Leach, 1815

Dyspessa hethitica Daniel, 1932

Dyspessa hethitica Daniel, 1932: Mitt. Münch. Ent. Ges 22: 16.

Type material examined

Syntype (ZMH 61343), 1♂ (Fig. 10). “CO-TYPE VON / DYSP/ HETHITICA DAN / F. DANIEL / MÜNCHEN // TAURUS / MARASCH / 6-900M a.-m.VII.31. / EINH SAMMLER LEG / F.DANIEL / MÜNCHEN // hethitica / Dan. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61343”.

Type locality

Taurus, Marasch [Turkey]

Current status

Valid species.

Remarks

Based on Daniel’s study (Daniel 1932), he labelled all specimens (1♂ and 5♀) of the type series as cotypes; therefore all specimens are syntypes. Holotype (by original designation) in Museum Witt München, MWM (Yakovlev 2011). ZMH used to hold two syntypes of D. hethitica (Weidner 1974); however, the female specimen could not be located any more.

Paropta frater (Warnecke, 1929)

Cossus frater Warnecke, 1929: Int Ent Z 23: 389.

Type material examined

Paratlectoype (ZMH 61344), 1♂ Südwest Arabien, Yemen, Sanäa, 2.viii.1931 (Fig. 11). “Paratypoid / SüdWest-Arabien / Yemen Samaa- / ? .Rathjens 2.8.31 // Cossus frater Warn. // Sig.G Warnecke / Eing Nr 5 1949 // ZMH 61344”.

Type locality

Arabien, Yemen, Sanäa.

Current status

Valid species.

Remarks

It was assumed that the syntypes were destroyed during the Allied bombing of Hamburg during World War II. However, as Warnecke reported to Daniel, two syntypes at the ZMH were lost in 1943, but one of them had been stored in Saxony and thus survived the bombings (Weidner 1974). Other type material (syntypes) is in the British Museum of Natural History (Yakovlev 2011; Yakovlev and Witt 2017).

Family Brachodidae Agenjo, 1966

Brachodes staudingeri Kallies, 1998

Brachodes staudingeri Kallies, 1998: Nota Lepidopterol. 21: 178.

Type material examined

Paratype (ZMH 61342), 1♂ (Fig. 12). “Asia centr. Kirgisia / 42’05,30N 75’06,96E / area W of Issyk Kul / Kys-Art Pass 2600m / 15. VII.1994 / Leg. A. Kallies // PARATYPUS / Brachodes / staudingeri sp. nov. / KALLIES, des. 1998 // ZMH 61342“.

Type locality

West of Issyk Kul Kys-Art Pass, Kirgizstan.

Current status

Valid species.

Remarks

Holotypus ♂, Kirgizstan, 42°05,30'N, 75°06,96'E, W of Issyk Kul, Kys-Art Pass, 2600 m, 15.vii.1994, leg. Kallies, Špatenka & Petersen (MfN – Museum für Naturkunde. Zentralinstitut der Humboldt-Universität. Berlin. Deutschland) (Kallies 1998).

Family Sesiidae Boisduval, 1828

Pyropteron chrysidiforme siculum f. fervens Bytinski-Salz, 1937

Pyropteron chrysidiforme siculum f. fervens Bytinski-Salz 1937: Mem. Soc. Ent. Ital. 15 (1936): 198.

Original material examined

Labelled as “Holotype” (ZMH 61326), 1♀ (Fig. 13); Labelled as “Paratype” 1♀ (ZMH 61327). “Oristano / Sardinia / 11/12.6.35. / Dr. H. Bytinski-Salz // chrysidi- / forme / fervens / ByS / f TYPE / ex coll. / Dr. H. Bytinski-Salz // Coll Bytinski-Salz / Eing Nr 20–60 // Pyropteron / chrysidiforme Esp. / f. ind. fervens / f Holotype / F. Le Cerf det. 1936 // ZMH 61326”; “chrysidi- / forme ab. / fervens / ByS / m PARATYPE / ex coll. / Dr. H. Bytinski-Salz // Pyr. chrysidifor / me. fervens / m (subtypique) / F. Le Cerf det. 1936 // Oristano / Sardinia / 11/12.6.35. / Dr. H. Bytinski-Salz // Coll Bytinski-Salz / Eing Nr 20, 1960 // ZMH 61327”.

Original locality

Sardinien, Oristano [Italy].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1937) proposed this name as a form of a Pyropteron chrysidiformesicilum . According to Article 45.6.4., the author expressly gave it infrasubspecific rank, in which case it is infrasubspecific, and hence unavailable.

Similipepsis osuni Bakowski & Kallies, 2008

Similipepsis osuni Bakowski & Kallies, 2008: Ann. Zool. 58 (4): 792.

Type material examined

Holotype (ZMH 61328), 1♂, W-Afrika, Nigeria, Ile-Ife, 13.v.1993 Ph. leg. L. Schlimm (Fig. 14). “Holotypus / Similipepsis / osuni / Bakowski & / Kallies 2008 // collection Riefenstahl / W-Afrika / Nigeria / Ile-Ife / 13.V.1993 Ph. / leg. L. Schlimm // ZMH 61328”.

Type locality

Ile-Ife, Osun State, Nigeria.

Current status

Valid species.

Sesia bembeciformis ab. incognita Strand, 1927

Sesia bembeciformis ab. incognita Strand, 1927: Arch. Naturg. 91 A (12): 282.

Original material examined

Labelled as “Holotype“ (ZMH 61329), 1♂, J. Androff, Hamburg, 18.viii.1911 (Fig. 15). „ZMH 61329 // Sig. G. Warnecke / Eing Nr 5. 1949 // J. Andorff / Hamburg / 18.8.11 // f. incognita Straude / Type / ? / Berlin, 1925, 9/ 12. / Abt.A / ? / 282“.

Original locality

Hamburg [Germany].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Strand (1927) proposed this name as an aberration of Sesia bembeciformis, therefore it is deemed to be infrasubspecific, which is hence unavailable (Article 45.6.2, ICZN 1999).

Bembecia abromeiti Kallies & Riefenstahl, 2000

Bembecia abromeiti Kallies & Riefenstahl 2000: Ent. Zs. 110 (12): 359.

Type material examined

Holotype (ZMH 828146), 1♂ (Fig. 16). “Holotypoid m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // Eingang: / ZMH 4/2001 // larva / pupa / 19.–22.V.2000 / ex Lotus cf. creticus // Espana, NE Mallorca / coast near Cala Canyamel, / S of Capdepera / 11.– 20. VI. 2000 e.l. / e.p. / lg. A. Kallies & A. Musolff // ZMH 828146“. Paratypes 18♂♀ (ZMH 61330–ZMH 61341, ZMH 828144–ZMH 828145, ZMH 828215–ZMH 828216, ZMH 833440–ZMH 833441) „PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // SPANIEN / MALLORCA / San Pedro / 39/44/50– 03/17/52 // 20.–30.5.2000 / leg. Riefenstahl/Roloff // ZMH 61330“; „SPANIEN / MALLORCA / San Pedro / 39/44/50–03/17/52 / 20.–30.5.2000 / leg. Riefenstahl/Roloff // PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61331“; „SPANIEN / MALLORCA / Calas de Mallorca / 39/28/02–03/16/14 / 20.–30.5.2000 / leg. Riefenstahl / Roloff // PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61332“; „SPANIEN / MALLORCA / Cala Agulla / 39/43/15–03/27/21 / 20.–30.5.2000 / leg. Riefenstahl / Roloff // PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61333“; „SPANIEN / MALLORCA / Cala Agulla / 39/43/15–03/27/21 / 20.–30.5.2000 / leg. Riefenstahl / Roloff // PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61334“; „SPANIEN / MALLORCA / San Pedro / 39/44/50–03/17/52 / 20.–30.5.2000 / leg. Riefenstahl/Roloff // PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61335“; „SPANIEN / MALLORCA / Cala Ratjada / 39/41/23–03/27/16 / leg. Riefenstahl / Roloff // PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61336“; „SPANIEN/MALLORCA / Cala Ratjada / 39/41/23–03/27/16 / e.p. 2.–15.6.2000 / leg. Riefenstahl / Roloff // PARATYPOID f / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61337“; „SPANIEN / MALLORCA / Cala Torta / 39/45/05–03/24/49 / 20.–30.5.2000 / leg. Riefenstahl / Roloff // PARATYPOID f / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61338“; „SPANIEN / MALLORCA / Cala Agulla / 39/43/15–03/27/21 / 20.–30.5.2000 / leg. Riefenstahl / Roloff // PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61339“; „SPANIEN / MALLORCA / Cala Ratjada / 39/41/23–03/27/16 / leg. Riefenstahl / Roloff // PARATYPOID f / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61340“; „SPANIEN / MALLORCA / Cala Ratjada / 39/41/23–03/27/16 / leg. Riefenstahl / Roloff // PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // ZMH 61341“; „SPANIEN / MALLORCA / Cala Agulla / 39/43/15–03/27/21 / 20.–30.5.2000 / leg. Riefenstahl / Roloff // PARATYPOID m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // Eingang: / ZMH 4/2001 // ZMH 828144“; „Paratypoid m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // SPANIEN / MALLORCA / Cala Agulla / 39/43/15–03/27/21 / 20–30.5.2000 / leg. Riefenstahl / Roloff // Eingang: / ZMH 4/2001 // ZMH 828145“; „Eing.-Nr. / ZMUH 1/2013 // Paratypoid m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // SPANIEN / MALLORCA / Cala Ratjada / 39/44/50–03/17/52 / 20.–30.5.2000 / leg. Riefenstahl / Roloff // ZMH 828215“; „Eing.-Nr. / ZMUH 1/2013 // Paratypoid m / Bembecia abromeiti / Kallies & / Riefenstahl / 2000 // SPANIEN / MALLORCA / Cala Ratjada / 39/41/23–03/27/16 / 20.–30.5.2000 / leg. Riefenstahl / Roloff // ZMH 828216”

Type locality

Espana, NE Mallorca [Spain].

Current status

Valid species.

Chamaesphecia (Scopulosphecia) minoica Bartsch & Puehringer, 2005

Chamaesphecia minoica Bartsch & Pühringer, 2005: Ent. Z. 115(3), 135.

Type material examined

Paratype (ZMH 828143), 1♂ (Fig. 17). “Paratypus / Chamaesphecia minoica / m / D. BARTSCH & / F. PÜHRINGER, DES. 2005 // Graecia, Kreta, 10–40 m / Marathi (Halbinsel Akrotiri) / 24,16°E / 35,51°N / 9.6.2000 // Pühringer & Pöll leg., Ph // Chamaesphecia / alysoniformis / (HERRICH-SCHÄFFER, 1846) / Pühringer det. 2000 // ZMH 828143”.

Type locality

Kreta, Graecia (Greece).

Current status

Valid species.

Family Metarbelidae Strand, 1909

Arbelodes claudiae Lehmann, 2010

Arbelodes claudiae Lehmann, 2010: 27–31.

Type material examined

Paratype (ZMH 835537), 1♀ (Fig. 18). “Arbelodes claudiae / I. Lehmann, 2010 / Paratype female, / North Malawi, Juniperus-Forest / 08.–11.October.2003 / R. J. Murphy leg. / Gen. slide number: / 19/03/2010 I. Lehmann // Def No 1193 / JUNIPER FOREST / NYIKA NAT PK N MALAWI / 2215m 1045S 3354E / 8–11 OCT 2003 R. J. Murphy // ZMH 835537”.

Type locality

Mzuzu, Nkhorongo, 1375 m., North Malawi.

Current status

Valid species.

Figures 10–18.

10. Dyspessa hethitica Daniel, 1932; 11. Paropta frater (Warnecke, 1929); 12. Brachodes staudingeri Kallies, 1998; 13. Pyropteron chrysidiforme siculum f. fervens Bytinski-Salz, 1937; 14. Similipepsis osuni Bakowski & Kallies, 2008; 15. Sesia bembeciformis ab. incognita Strand, 1927; 16. Bembecia abromeiti Kallies & Riefenstahl, 2000; 17. Chamaesphecia (Scopulosphecia) minoica Bartssch & Puhringer, 2005; 18. Arbelodes claudiae Lehmann, 2010.

Superfamily Zygaenoidea Latreille, 1809

The definition of the superfamily Zygaenoidea is based on only few characters. Although these characters are possibly autapomorphies this is partly not confirmed and is still a matter of discussion. The discussed characters are: head retractibility in larva, heteromorphosis, forward migration and reduction of head setae, position of microsetae and pores, labral sensilla (Vegliante and Zilli 2004). However, most of these characters have still not been sufficiently studied throughout all families. An overview regarding this discussion can be found in Efetov and Tarmann (2017). The Zygaenoidea contain the 13 families: Epipyropidae, Cyclotornidae, Himantopteridae, Anomoeotidae, Megalopygidae, Somabrachidae, Aididae, Limacodidae, Dalceridae, Lacturidae, Heterogynidae, Phaudidae, and Zygaenidae (Epstein et al. 1999; Efetov et al. 2014).

The family Zygaenidae has a world-wide distribution and is divided into the five subfamilies Inouelinae, Procridinae, Chalcosiinae, Callizygaeninae and Zygaeninae (Efetov and Tarmann 2017); it includes more than 1,000 species. 25 relevant characters that are possibly part of a ‘hypothetical groundplan’ of the family are discussed in detail in Efetov and Tarmann (2017). The most relevant possible autapomorphies of Zygaenidae are: 1. Cyanogenesis and resistance against cyanides. 2. Head with ocelli and chaetosemata present (as in some other more primitive ditrysian groups, e.g. Yponomeutoidea and Tortricoidea). This character combination is absent in all other Zygaenoidea. 3. Development of Petersen’s gland (a pair of glands close to the ooporus in Zygaeninae and Procridinae of not exactly known function). This character is considered to be secondarily reduced in Inouelinae, Chalcosiinae and Callizygaeninae. This gland seems to have an importance for the protection of the eggs against predators.

Family Zygaenidae Latreille, 1809

Subfamily Zygaeninae Latreille, 1809

Zygaena (Agrumenia) carniolica berolinensis v. pinskica Reiss, 1941

Zygaena (Agrumenia) carniolica Scop. subsp. berolinensis Stgr. var. pinskica n. var. (subsp. ?): Z. Wien. Ent. Ver. 26: 59.

Original material examined

Labelled as “Cotype” 5♂♀ (ZMH 61362−61365, 61367) (Fig. 19). “Piokitao / Sümpfe / 18.7.17. // // Cotype f / pinskica / Reiß 1941. / H.Reiss // ZMH 61362”; “Rokituo / Sümpfe / 17.7.17. // // Cotype m / pinskica / Reiß 1941. / H.Reiss // ZMH 61363”; “Rokituo / Sümpfe / 18.7.17. // // Cotype m / pinskica / Reiß 1941. / H.Reiss // ZMH 61364”; “Rokituo / Sümpfe / 17.7.17. // // Cotype f / pinskica / Reiß 1941. / H.Reiss // ZMH 61365”; “Rofrituo / Sümpfe / 12.7.17. // // Cotype m / pinskica / Reiß 1941. / H.Reiss // ZMH 61367”.

Original locality

‘Pinsk, Lesnaja (Rokitno)’ [Belarus: Pinsk, Ljesnaja (Roknito)] (Hofmann and Tremewan 1996: 112).

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1941) proposed this name as a local variant of Z. c. berolinensis Lederer, 1853 with the remark ‘var. pinskica n. var. (subsp.?)’. According to Article 45.6.4., the author expressly gave it infrasubspecific rank, in which case it is infrasubspecific, and hence unavailable.

Zygaena (Agrumenia) carniolica pinskica ab.cingulata Reiss, 1941

Zygaena (Agrumenia) carniolica pinskica ab. cingulata Reiss, 1941: Z. Wien. Ent. Ver. 26: 59.

Original material examined

Labelled as “Type” 1♂ (ZMH 61366) (Fig. 20). “Rokinto / Sümpfe / 10.7.17. // // Type // ab. cingulata Reiss // Type m / 1941 / pinskica / Reiß ab. / cingulata / Reiß H. Reiss // ZMH 61366”.

Original locality

‘Pinsk, Lesnaja (Rokitno)’ [Belarus: Pinsk, Ljesnaja (Roknito)] (Hofmann and Tremewan 1996: 112).

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1941) proposed this name as an aberration of Z. c. pinskica Reiss, 1941. Therefore, as stated by Articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be infrasubspecific name – if author used “aberration”, “ab.”, “morph”, or the author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena (Zygaena) angelicae ternovanensis Koch, 1938

Zygaena (Zygaena) angelicae ternovanensis Koch, 1938: Z. Öst. Ent. Ver. 23: 17–18.

Type material examined

Syntypes 3♂♂ (ZMH 61368−61370) (Fig. 21). “? / 7.1932 // v. ternovanensis / Koch // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61368”; “? / e.l. 6.1927. / P. Ronnicke // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61369”; “Tamooamer / Wald // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61370”.

Type localities

Slovenia [Julisches Venetien]: Nova Gorizia [Görz], Trnovski Gozd [Ternovaner-Wald] (Hofmann and Tremewan 1996)

Current status

Valid subspecies

Zygaena (Zygaena) angelicae ternovanensis ab. ternoflava Koch, 1938

Zygaena (Zygaena) angelicae ternovanensis ab.ternoflava Koch, 1938: Z. Öst. Ent. Ver. 23: 17–18.

Original material examined

Labelled as “Cotype” 2♂♂ (ZHM 61371– ZMH 61372) (Fig. 22). „Ternov. nes / 7.1932 // Cotypus // Original- / material / der Zuchten / von / Ronnicker / Graz // ab. ternoflava / Koch // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61371“; „? / 7.1932 // Cotypus / Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61372“

Original localitiy

‘Jugoslawien, Julisches Venetien, Ternovaner Wald, nordöstlich von Görz, 800–900 m.‘ [Slovenia: Nova Gorizia, Trnovski Gozd]

Current status

Infrasubspecific and hence unavailable name.

Remarks

Koch (1938) proposed this name as an aberration of Z. c. ternovanensis Koch, 1938, therefore as stated by Articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be infrasubspecific name – if author used “aberration”, “ab.”, “morph”, or the author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena (Mesembrynus) corsica sardiniensis ab. minor Bytinski-Salz, 1937

Zygaena corsica sardiniensis ab.minor Bytinski-Salz, 1937: Mem. Soc. Ent. Ital. 15 (1936): 196.

Original material examined

Labelled as “Type” 1♂ (ZHM 61374) (Fig. 23). “Aritzo / Sardinia / 900 m / 10/22.6.1934 / Dr. H. Bytinski-Salz // Zyg. corsica / wp-Sardinia / ? / ab. minor / ByS / m TYPE / ex.coll. / Dr. H. Bytinski-Salz // ab. minor / By.S. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61374”.

Original locality

Aritzo, Sardinia [Italy].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1937) proposed this name as an aberration of Z. c. sardiniensis Holik, 1936, therefore as stated by Articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be infrasubspecific name – if author used “aberration”, “ab.”, “morph”, or the author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena (Mesembrynus) corsica sardiniensis ab. bielongata Bytinski-Salz, 1937

Zygaena corsica sardiniensis ab.bielongata Bytinski-Salz, 1937: Mem. Soc. Ent. Ital. 15 (1936): 196.

Original material examined

Labelled as “Type” 1♂ (ZHM 61375) (Fig. 24). “Aritzo / Sardinia / 900 m / 10/22.6.1934 / Dr. H. Bytinski-Salz // corsica / ab. / bielongata / Bys. / TYPE / ex. coll. / Dr. H. Bytinski-Salz // ab. bielongata / ByS. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61375”.

Original locality

Aritzo, Sardinia [Italy].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1937) proposed this name as an aberration of Z. c. sardiniensis Holik, 1936. Therefore, as stated by Articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be infrasubspecific name – if author used “aberration”, “ab.”, “morph”, or the author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena (Agrumenia) cocandica banghaasi Burgeff, 1927

Zygaena (Agrumenia) cocandica banghaasi Burgeff, 1927: Horae Macrolep. Reg. Pal. 1: xxvii: 56.

Type material examined

Syntypes 2♂♂ (ZHM 61376– ZMH 61377) (Fig. 25). “Dombratschi / Buchara merid. / 28.Juli 2000m. // Cotypus / cocandica / v. BangHaasi / Bgff. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // v. banghaasi / Bgff. // ZMH 61376 “; “Dombratschi / Buchara merid. / 28.Juli 2000m. // Cotypus / cocandica / v. BangHaasi / Bgff. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61377”.

Type locality

‘Buchara, Karategin mont. mer. or.’ [Tajikistan: Karategin Range].

Current status

Synonym of Zygaena (Agrumenia) cocandica cocandica Erschoff, 1874.

Remarks

Burgeff (1927) proposed this name as subspecies of Z. cocandica, but this taxon was synonymized with Zygaena (Agrumenia) cocandica cocandica Erschoff, 1874 (Hofmann and Tremewan 1996).

Zygaena (Mesembrynus) purpuralis chamurli Koch, 1935

Zygaena purpuralis ssp. chamurli Koch, 1935 (1934): Dt. Ent. Z. Iris 48 (1934): 192.

Type material examined

Paralectotypes 4♂♂. (ZHM 61378– ZMH 61381) (Fig. 26). «Kurdistan / Chanuria-Dagh / Geröllzone / 2900m 19–21.7. / leg. Kotzsch // chamurli / Koch // Cotype // 621. // Coll. Bytinski-Salz Coll. / Eing. Nr. 20, 1960 Eing. // ZMH 61378»; «Kurdistan / Chanuria-Dagh / Geröllzone / 2900m 19–21.7. / leg. Kotzsch // Cotype // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // 618. // ZMH 61379”; “Kurdistan / Chanuria-Dagh / Geröllzone / 2900m 19–21.7. / leg. Kotzsch // Cotype // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // 619. // ZMH 61380”; “Kurdistan / Chanuria-Dagh / Geröllzone / 2900m 19–21.7. / leg. Kotzsch // Cotype // 620. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61381”.

Type locality

‘Kurdistan. Chamurlu-Dagh, bei Sarikamisch, 70 km südwestlich von Kars, Geröllhalden, 2900 m, 19–21.xii.1934‘[NE Turkey: Kars, Sarikamiş, Çamurlu Dağı].

Current status

Valid subspecies. Under revision.

Remarks

Based on the revision of Naumann (1982), he designated lectotype from an original specimen described by Koch from the collection of “Staatl. Museum für Tierkunde Dresden“. Therefore, the rest of the type specimens are paralectotypes. The taxon group containing Zygaena purpuralis chamurli Koch, 1935 is currently revised by Ana Nahirnić (National Museum of Natural History, Bulgaria, Sofia).

Zygaena (Mesembrynus) contaminei asturica Reiss, 1936

Zygaena (Mesembrynus) contaminei asturica Reiss, 1936: Entomologische Rundschau 54: 59.

Type material examined

Syntypes 2♂♂ (ZHM 61382– ZMH 61383) (Fig. 27). “Picos Europa / da Liebana / A. Kricheldorff // Cotypus / Z. contaminei / ssp. / asturica / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ssp. asturica / Reiß. // ZMH 61382”; “Picos Europa / da Liebana / A. Kricheldorff // Cotypus / Z. contaminei / ssp. / asturica / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61383”.

Type locality

‘La Liébana’ [Spain: Cantabria, La Vega de Liébana] (Hofmann and Tremewan 1996).

Current status

Synonym of Zygaena contaminei penalabrica Fernández, 1929.

Remarks

Synonym to Zygaena contaminei penalabrica Fernández, 1929 (Hofmann and Tremewan 1996).

Zygaena (Mesembrynus) corycia r. wiltshirei Bytinski-Salz, 1936

Zygaena corycia ‘race’ wiltshirei Bytinski-Salz, 1936: Ent. Rec. J. Var. 48 (Suppl.): (1)–(6): (1).

Original material examined

Labelled as “Paratype” 1♂ (ZHM 61384) (Fig. 28). “SYRIA // Z. corycia / ssp. / wiltshirei / By.S. / PARATYPE / ex coll. / Dr. H. Bytinski-Salz // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ssp. wiltshirei / By.S. // ZMH 61384”.

Original locality

‘Kineseh’ [Lebanon: Djebel Kineseh].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1936) proposed this name as a ‘race’ of Z. corycia Staudinger, 1871. Therefore, as stated by Article 45.6.1 (“the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity”) it is deemed to be infrasubspecific name, which is hence unavailable. Zygaena corycia was downgraded to a subspecies of Z. (Mesembrynus) brizae (Esper, 1800) (Hofmann and Tremewan 1996).

Figures 19–28.

19. Zygaena (Agrumenia) carniolica v. pinskica Reiss, 1941; 20. Zygaena (Agrumenia) carniolica pinskica ab. cingulata Reiss, 1941; 21. Zygaena (Zygaena) angelicae ternovanensis Koch, 1938; 22. Zygaena (Zygaena) angelicae ternovanensis ab. ternoflava Koch, 1938; 23. Zygaena (Mesembrynus) corsica sardiniensis ab. minor Bytinski-Salz, 1937; 24. Zygaena (Mesembrynus) corsica sardiniensis ab. bielongata Bytinski-Salz, 1937; 25. Zygaena (Agrumenia) cocandica banghaasi Burgeff, 1927; 26. Zygaena (Mesembrynus) purpuralis chamurli Koch, 1935; 27. Zygaena (Mesembrynus) contaminei asturica Reiss, 1936; 28. Zygaena (Mesembrynus) corycia r. wiltshirei Bytinski-Salz, 1936.

Zygaena (Mesembrynus) brizae staudingeriana Reiss, 1932

Zygaena (Mesembrynus) corycia staudingeriana Reiss, 1932: Int. Ent. Z. 26: 270.

Type material examined

Syntypes 6♂♂ (ZHM 61385– ZMH 61390) (Fig. 29). “SYRIA / LIBANON / BSCHARRE / 1600M 1,–15 VI.31 / E. PFEIFFER LEG / F DANIEL / MÜNCHEN // CO-TYPE VON / Z. CORYCIA SSP. / STAUDINGERIANA / REISS / DANIEL / MÜNCHEN // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // v. staudingeralis / Reiß. // ZMH 61385”; “SYRIA / LIBANON / BSCHARRE / 1600M 1,–15 VI.31 / E. PFEIFFER LEG / F DANIEL / MÜNCHEN // CO-TYPE VON / Z. CORYCIA SSP. / STAUDINGERIANA / REISS / DANIEL / MÜNCHEN // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61386”; “SYRIA / LIBANON / BSCHARRE / 1300M 1, –15 VI.31 / E. PFEIFFER LEG / F DANIEL / MÜNCHEN // CO-TYPE VON / Z. CORYCIA SSP. / STAUDINGERIANA / REISS / DANIEL / MÜNCHEN // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61387”; “Becharree / Libanon / Juni 31. / Kulzer. // CO-TYPE VON / Z. CORYCIA SPP. / STAUDINGERIANA / REISS / DANIEL / MÜNCHEN // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61388”; “Becharree / Libanon / Juni 31. / Kulzer // CO-TYPE VON / Z. CORYCIA SPP. / STAUDINGERIANA / REISS / DANIEL / MÜNCHEN // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61389”; “Becharree / Libanon / Juni 31. / Kulzer // CO-TYPE VON / Z. CORYCIA SPP. / STAUDINGERIANA / REISS / DANIEL / MÜNCHEN // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61390”.

Type locality

Libanon, Bscharre [Lebanon].

Current status

Valid subspecies.

Remarks

Reiss (1932a) proposed this name as a subspecies of Z. corycia Staudinger, 1871, but this species name was synonymized with Z. (Mesembrynus) brizae (Esper, 1800) (Hofmann and Tremewan 1996), thus the current combination is Z. (M.) brizae staudingeriana Reiss, 1932.

Zygaena (Mesembrynus) corycia r. amseli Bytinski-Salz, 1936

Zygaena corycia ‘race’ amseli Bytinski-Salz, 1936: Ent. Rec. J. Var. 48 (Suppl.): (1)–(6): 2.

Original material examined

Labelled as “Cotype” 4♂♂ (ZHM 61391– ZMH 61394) (Fig. 30). “Ain Karem / b. Jerusalem / 21.4.30 / leg. H. G. Amsel // Cotypus / Corycia / ssp. / amseli / By.S. // ssp. amseli / ByS. // Corycia / Stgr. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61391“; „Ain Karem / b. Jerusalem / 21.4.30 / leg. H. G. Amsel // Cotypus / Corycia / ssp. / amseli / ByS. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61392”; “Ain Karem / b. Jerusalem / 21.4.30 / leg. H. G. Amsel // Cotypus / Corycia / ssp. / amseli / By.S. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61393“; „Ain Karem / b. Jerusalem / 21.4.30 / leg. H. G. Amsel // Cotypus / Corycia / ssp. / amseli / ByS. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61394”.

Original locality

‘Ain Karem, Palestine’ [Israel: Jerusalem West, En Kerem].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Bytinski-Salz (1936) proposed this name as ‘race’ (= subspecies) of Z. corycia Staudinger, 1871. Therefore, as stated by Article 45.6.1 (“the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity”) it is deemed to be infrasubspecific name, which is hence unavailable.

Zygaena (Zygaena) ephialtes ab. maureri Dziurzynski, 1913

Zygaena ephialtesab.maureri Dziurzynski 1913: Jber. Wien. Ent. Ver. 23 (1912): 215.

Original material examined

Labelled as “Type” 1♂ (ZMH 61395) (Fig. 31). „Weiz oder Wuz / 22/7 // ex coll. Ronnicke / TYPE // ab. maureri / DZ. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61395“.

Original locality

‘Steiermark, Weiz (in der Raaberklamm)‘ [Austria: Styria, Weiz, Rabenklamm].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Dziurzynski (1913) proposed this name as an aberration of Z. ephialtes (Linnaeus, 1767). Therefore, it is deemed to be infrasubspecific, it is hence unavailable (Article 45.6.2, ICZN 1999).

Zygaena (Zygaena) ephialtes kiewensis Reiss, 1932

Zygaena ephialtes kiewensis Reiss, 1932: Dt. Ent. Z. Iris 26: 130.

Type material examined

Syntype 1♂ (ZHM 61396) (Fig. 32). “ ? / 9.VII.30 // Kiev // Cotype m / kiewensis / Reiß 1932 / H. Reiss // kiewensis / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61396”.

Type locality

‘Ukraine: Kirillovskije ovragi’ [Ukraine: vicinity of Kiyev].

Current status

Synonym of Z. ephialtes athamanthae (Esper, 1789).

Remarks

Reiss (1932b) in Holik (1932) proposed this name as a subspecies of Z. ephialtes (Linnaeus, 1767). The name was subsequently synonymized with Z. ephialtes athamanthae (Esper, 1789) (Hofmann and Tremewan 1996).

Zygaena (Zygaena) dorycnii korbiana Reiss, 1935

Zygaena dorycnii korbiana Reiss, 1935: Int. Ent. Z. 29: 230

Type material examined

Syntype 1♂ (ZHM 61400) (Fig. 33). “Achalzich / (Chamoobel) / 1910 Korb // ssp. korbiana / Reiß // Cotypus / dorycnii / ssp. korbi- / ana / Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61400”.

Type locality

Transkaukasien, Akhaltsikhe [Georgia, Chambobel].

Current status

Synonym of Z. dorycnii dorycnii Ochsenheimer, 1808.

Remarks

Reiss (1935c) proposed this name as subspecies of Z. dorycnii Ochsenheimer, 1808. The name was subsequently synonymized with the nominotypical Z. dorycnii dorycnii Ochsenheimer, 1808 (Hofmann and Tremewan 1996).

Zygaena (Agrumenia) faustaagilis Reiss, 1932

Zygaena fausta agilis Reiss, 1932: Int. Ent. Z. 29: 227.

Type material examined

Syntypes 2♂♂ (ZHM 61401– ZMH 61402) (Fig. 34). “Zyg. fausta / Jena 10.72 // Cotype m / agilis / Reiß / H. Reiss // ssp. agilis / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61401”; “Zyg. fausta / Jena 10.72 // Cotype f / agilis / Reiß / H. Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61402”.

Type locality

Thüringen, Jena [Germany].

Current status

Valid subspecies.

Remarks

Reiss (1932c) proposed this name as subspecies of Z. fausta (Linnaeus, 1767) that is considered a valid subspecies name.

Zygaena (Agrumenia) exulans sajana Burgeff, 1926

Zygaena exulans sajana Burgeff, 1926: Mitteilungen der Münchner Entomologischen Gesellschaft 16: 25.

Type material examined

Paratypes 3♂♂ (ZHM 61403– ZMH 61405) (Fig. 35). “Munko Sardyk / Sajan mont. // Cotypus / exulans / ssp. sajana / Bgff // ssp. sajana / Bgff. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61403”; “Schawyr / Tannuola or. / Juni 2500 m // Cotypus / exulans / ssp. sajana / Bgff // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61404”; “Schawyr / Tannuola or. / Juni 2500m // Cotypus / exulans / ssp. sajana / Bgff // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61405”.

Type localities

‘Sajangebirge (Munko Sardyk und Schawyr, Tannuola or.)‘ [Munko-Sardyk mountain in Western Buryatia near Mongolian Border].

Current status

Valid subspecies.

Remarks

Burgeff (1926) proposed this name as a subspecies of Z. exulans (Hohenwarth, 1792) that is considered a valid subspecies name. As Burgeff’s holotypes are in the Eidgenössische Technische Hochschule (ETH), Zürich, in Switzerland, these cotypes must be paratypes.

Zygaena (Agrumenia) exulans polaris Holik, 1935

Zygaena exulans var. polaris Holik, 1935: Dalm. Ent Tisdkr 56, 47.

Type material examined

Syntypes 2♂♂ (ZHM 61406– ZMH 61407) (Fig. 36). «Rybatschi H.-Ins / Murman-Küste / Fjeldzone 300m. / 70° N.Br. 28.6.–8.7 / leg. Kotzsch // Cotypus / Z. exulans / ssp. polaris / Holik // ssp. polaris / Holik. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61406“; „Rybatschi H.-Ins / Murman-Küste / Fjeldzone 300 m. / 70° N. Br. 28.6.–8.7 / leg. Kotzsch // Cotypus / Z. exulans / ssp. polaris / Holik // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61407”.

Type locality

Rybatschiy H.-Ins, Murman-Küste, Fjeidzone, 150 m.

Current status

Synonym of Z. exulans vanadis Dalman, 1816.

Remarks

Holik (1935) proposed this name as a variation of Z. exulans (Hohenwarth, 1792) from Arctic Russia (Murmansk area). According to Article 45.6.4., it is subspecific if first published before 1961 and author expressly used the terms “variety”. This ‘variation’ was a group of populations with a defined range and was therefore treated as a subspecies by Reiss and Tremewan (1967). Later, Hofmann and Tremewan (1996) synonymised it with Z. exulans vanadis Dalman, 1816. ‘variatio’, in 1935 this term was often used in the current sense of subspecies.

Figures 29–36.

29. Zygaena (Mesembrynus) brizae staudingeriana Reiss, 1932; 30. Zygaena (Mesembrynus) corycia r. amseli Bytinski-Salz, 1936; 31. Zygaena (Zygaena) ephialtes ab. maureri Dziurzynski, 1913; 32. Zygaena (Zygaena) ephialtes kiewensis Reiss, 1932; 33. Zygaena (Zygaena) dorycnii korbiana Reiss, 1935; 34. Zygaena (Agrumenia) fausta agilis Reiss, 1932; 35. Zygaena (Agrumenia) exulans sajana Burgeff, 1926; 36. Zygaena (Agrumenia) exulans polaris Holik, 1935.

Zygaena (Mesembrynus) graslini rebeli Reiss, 1932

Zygaena graslini Lederer subsp. rebeli Reiss, 1932: Int. Ent. Z. 26: 275.

Type material examined

Syntypes 6♂♀ (ZHM 61412– ZMH 61417) (Fig. 37). “AMANUS S. / DÜL-DÜL DAGH / YÜKSEK DAGH / ANF.-M. IV 32. / EINH. SAMMLER LEG. / F DANIEL / MÜNCHEN // Cotypus // ssp. rebeli / Reiß. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61412“; „AMANUS S. / DUL-DUL DAGH / YUKSEK DAGH / ANF.-M. IV 32. / EINH SAMMLER LEG. / F. DANIEL / MÜNCHEN // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61413“; „AMANUS S. / DUL-DUL DAGH / YUKSEK DAGH / ANF.-M. IV 32. / EINH SAMMLER LEG. / F. DANIEL / MÜNCHEN // CO-TYPE VON / Z. GRASLINI / SSP. REBELI / REISS / DANIEL / MÜNCHEN // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61414“; „AMANUS S. / DUL-DUL DAGH / YUKSEK DAGH / ANF.-M. IV 32. / EINH SAMMLER LEG. / F. DANIEL / MÜNCHEN // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61415“; „AMANUS S. / DUL-DUL DAGH / YUKSEK DAGH / ANF.-M. IV 32. / EINH SAMMLER LEG. / F. DANIEL / MUNCHEN // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61416“; „AMANUS S. / DUL-DUL DAGH / YUKSEK DAGH / ANF.-M. IV 32. / EINH SAMMLER LEG. / F. DANIEL / MÜNCHEN // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61417“.

Type locality

Yüksek Dagh, Amanus.

Current status

Invalid subspecies. There are currently no subspecies accepted in Zygaena graslini. Synonym of Zygaena (Mesembrynus) graslini s.str.

Zygaena (Mesembrynus) hindukuschi Koch, 1937

Zygaena hindukuschi Koch, 1937: Entomologische Zeitschrift. Frankfurt a. M. 51: 71.

Type material examined

Syntype 1♂ (ZMH 61460) (Fig. 38). „Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H. & E.Kotzsch // hindukuschi / Koch. // Cotypus / Zygaena / hindukuschi / Koch // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61460”.

Type locality

Afghanistan (Badakhshan)

Current status

Valid species.

Zygaena (Agrumenia) hilaris lucifera Reiss, 1936

Zygaena hilaris ssp. lucifera Reiss, 1936: Entomologische Rundschau 54: 72.

Type material examined

Syntypes 6♂♀ (ZMH 61461– ZMH 61466) (Fig. 39). «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 61461»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 61462»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 61463»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 61464»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 61465»; «Cotypen // Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 61466».

Type locality

‘Sierren um Albarracin’ [Spain: Teruel, Sierra de Albarracin]

Current status

Valid subspecies.

Zygaena (Agrumenia) hilaris lucifera ab. pseudocatalonica Reiss, 1936

Zygaena hilaris ssp. lucifera ab.pseudocatalonica Reiss, 1936: Entomologische Rundschau 54: 73.

Original material examined

Labelled as “Cotype” 1♂1♀ (ZMH 61467, ZMH 61470) (Fig. 40). «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Cotype // 1 f Cotype lucifera / ab. pseudocataloni- / ca Reiß 1936 // ZMH 61467“; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ab. pseudo- / catalonica Reiss // Type // Type m / lucifera Reiß / ab. pseudoca- / talonica Reiß / 1936 H.Reiss // ZMH 61470”.

Original locality

‘Sierren um Albarracin’ [Spain: Teruel, Sierra de Albarracin]

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1936) proposed this name as an aberration of Z. h. lucifera Reiss, 1936. Therefore, as stated by Articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be infrasubspecific name – if author used “aberration”, “ab.”, “morph”, or the author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena (Agrumenia) hilaris lucifera ab. inversa Reiss, 1936

Zygaena hilaris ssp. lucifera ab.inversa Reiss, 1936: Entomologische Rundschau 54: 73.

Original material examined

Labelled as “Type” 2♂♀ (ZMH 61468– ZMH 61469) (Fig. 41). “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ab. inversa Reiss // Type // Type m / lucifera Reiß / ab. inversa / Reiß / 1936. H.Reiss // ZMH 61468”; “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type f / lucifera Reiß / ab. inversa / Reiß / 1936 H.Reiss // ZMH 61469”.

Original locality

‘Sierren um Albarracin’ [Spain: Teruel, Sierra de Albarracin]

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1936) proposed this name as an aberration of Z. h. lucifera Reiss, 1936. Therefore, as stated by Articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be infrasubspecific name – if author used “aberration”, “ab.”, “morph”, or the author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena (Agrumenia) hilaris lucifera ab. gelpkei Reiss, 1936

Zygaena hilaris ssp. lucifera ab.gelpkei Reiss, 1936: Entomologische Rundschau 54: 73.

Original material examined

Labelled as “Cotype” 1♀ (ZMH 61471) (Fig. 42). “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type f / lucifera Reiß / ab. gelpkei / Reiß / 1936 H.Reiss // ZMH 61471”.

Original locality

No type-locality mentioned in original description. This type comes from Sierra Alta de Albarracin.

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1936) proposed this name as an aberration of Z. hilaris lucifera Reiss, 1936. Therefore, as stated by Articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be infrasubspecific name – if author used “aberration”, “ab.”, “morph”, or the author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena (Agrumenia) hilaris lucifera ab. exarcuata Reiss, 1936

Zygaena (Agrumenia) hilaris lucifera ab.exarcuata Reiss, 1936: Entomologische Rundschau 54: 73.

Original material examined

Labelled as “Cotype” 1♂ (ZMH 61472) (Fig. 43). “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type m / lucifera Reiß / ab. exarcua – / ta / Reiß / 1936 H.Reiss // ZMH 61472”.

Original locality

No type locality given for this taxon. This type comes from Sierra Alta de Albarracin.

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1936) proposed this name as an aberration of Z. hilaris lucifera Reiss, 1936. Therefore, as stated by Articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be infrasubspecific name – if author used “aberration”, “ab.”, “morph”, or the author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena (Mesembrynus) laeta danieli Reiss, 1935

Zygaena laeta danieli Reiss, 1935: Int. Ent. Z. 29: 159.

Type material examined

Syntype 1♂ (ZMH 61473) (Fig. 44). “Asia min.c. / Ak-Chehir / 21.–30.Juni 28 / Coll.Wagner-Wien // Cotypus / ssp. danieli / Reiß // ssp. danieli / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61473”.

Type locality

‘Marasch’ [Turkey: Kahraman Maraş]

Current status

Synonym of Z. laeta akshehirensis Reiss, 1929.

Remarks

Reiss (1935c) proposed this name as subspecies of Zygaena laeta (Hübner, 1790). The name was subsequently synonymized with current combination Z. laeta akshehirensis Reiss, 1929 (Hofmann and Tremewan 1996).

Zygaena (Agrumenia) formosa kotzschi Reiss, 1935

Zygaena (Agrumenia) formosa kotzschi Reiss, 1935: Int. Ent. Z. 28: 489.

Type material examined

Syntypes 3♂♂ (ZMH 61475– ZMH 61477) (Fig. 45). „Cotype m / kotzschi / Reiß 1935 / H. Reiss // Armenien / Rharhkhash-Dagh / Geröll-Nordabbang / 3200m. 1.–10 7. / leg. Kotzsch // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // 623. // ZMH 61475“; „Armenien / Rharhkhash-Dagh / Geröll-Südabhang / 3200m. 1.–10 7. / leg. Kotzsch // Cotype // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // 624. // ZMH 61476“; „Cotype m / kotzschi / Reiß 1935 / H. Reiss // Armenien / Rharhkhash-Dagh / Geröll-Südabhang / 3200m. 1.–10 7. / leg. Kotzsch // kotzschi / Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // 622. // ZMH 61477“.

Type locality

‘Armenien, Khashkhash-Dagh’ [Turkey: Kars, Ağrı Dağ, Kash-kash Dağ].

Current status

Valid subspecies.

Zygaena (Mesembrynus) araxis Koch, 1936

Zygaena manlia ssp. araxis Koch, 1936: Dt. Ent. Z. Iris 50: 41.

Type material examined

Syntypes 2♂♂ (ZMH 61478– ZMH 61479) (Fig. 46). “Armenia ross. / Ordubad / 900 m. // Co-Type / e Collection / Bang-Haas // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ssp. araxis (written on label as ‘aiaxis’ [sic]) / Koch // ZMH 61478”; “Armenia ross. / Ordubad / 900 m. // Co-Type // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61479”.

Type locality

Nus.nus bei Ordubad‘ [Azerbaijan: Nakhichevan, near Orduba, Nyusnyus (Nus-nus)] (Hofmann and Tremewan 1996).

Current status

Valid species.

Remarks

Zygaena araxis is a valid species. According to Hofmann and Tremewan (2020) its taxonomic status has a confusing history. It was raised to species level by Görgner and Hofmann (1982), but subsequently placed as a subspecies to Z. rubricollis Hampson, 1900 by Hofmann and Tremewan (1996). Finally, it was reinstated as a valid species by Keil (2003).

Figures 37–46.

37. Zygaena (Mesembrynus) graslini rebeli Reiss, 1932; 38. Zygaena (Mesembrynus) hindukuschi Koch, 1937; 39. Zygaena (Agrumenia) hilaris lucifera Reiss, 1936; 40. Zygaena (Agrumenia) hilaris lucifera ab. pseudocatalonica Reiss, 1936; 41. Zygaena (Agrumenia) hilaris lucifera ab. inversa Reiss, 1936; 42. Zygaena (Agrumenia) hilaris lucifera ab. gelpkei Reiss, 1936; 43. Zygaena (Agrumenia) hilaris lucifera ab. exarcuata Reiss, 1936; 44. Zygaena (Mesembrynus) laeta danieli Reiss, 1935; 45. Zygaena (Agrumenia) formosa kotzschi Reiss, 1935; 46. Zygaena (Mesembrynus) araxis Koch, 1936

Zygaena (Agrumenia) truchmena magnifica Reiss, 1935

Zygaena magnifica Reiss 1935: Int. Ent. Z. 29: 41.

Type material examined

Syntypes 6♂♀ (ZMH 61480– ZMH 61485) (Fig. 47). “Buchara / Kurgantjube // Cotypus / Z. magnifica / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61480”; “Buchara / Kurgantjube // Cotypus / Z. / magnifica / Reiß // Coll. Warnecke / Eing. Nr. 5, 1949 // ZMH 61481”; “Kurgan Tjube / Buchara / 7.1934 // Cotype f / magnifica / Reiß / 1935 H.Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61482”. Kurgan Tjube / Buchara / Juli 1934. // Cotype m / magnifica / Reiß 1935 / H.Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61483”; “Kurgan Tjube / Buchara / 7.1934 // Cotype m / magnifica / Reiß / 1935 H. Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61484”. “Kurgan Tjube / Buckara / 7. 1934 // Cotype m / magnifica / Reiß / 1935. H. Reiss // magnifica / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61485”.

Type localitiy

Kurgan-Tyube [Tajikistan].

Current status

Valid subspecies.

Remarks

Downgraded to subspecific rank. The current combination is Z. truchmena magnifica Reiss 1935 (Hofmann and Tremewan 1996).

Zygaena (Zygaena) lonicerae stettinensis Reiss, 1922

Zygaena lonicerae stettinensis Reiss, 1922: Int. Ent. Z. 16: 66.

Type material examined

Syntypes 1♂1♀ (ZMH 61486–ZMH 61487) (Fig. 48). “Hoekendorf / H. Rüffer. / 2.8.22. // Cotype f / stettinensis / Reiß / 1922 H. Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61486”. “Hoekendorf / H. Rüffer. / 5.8.22. // v. stettinensis / Reiss // Cotype m / stettinensis / Reiss / 1922 H.Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61487”.

Type locality

Stettin [Poland].

Current status

Synonym of Z. lonicerae lonicerae (Scheven, 1777).

Remarks

Reiss (1922) proposed this name as a subspecies of Z. lonicerae (Scheven, 1777), but this subspecific name was synonymized with Z. lonicerae lonicerae (Scheven, 1777) (Hofmann and Tremewan 1996).

Zygaena (Zygaena) lavandulae barcelonica Reiss, 1936

Zygaena lavandulae barcelonica Reiss, 1936: Entomologische Rundschau 54: 71.

Type material examined

Syntypes 1♂1♀ (ZMH 61488–ZMH 61489) (Fig. 49). “? / Cotype m / barcelonica / Reiß / 1936 H. Reiss // ssp. barcelonica / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61488”; “Barcelona / a. ? // Cotype f / barcelonica / Reiß / 1936. H.Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61489”.

Type locality

Barcelona [Spain].

Current status

Valid subspecies.

Zygaena (Zygaena) meliloti silaecola Verity, 1930

Zygaena meliloti silaecola Verity, 1930: Memorie della Societa Entomologica Italiana 9: 12.

Original material examined

“Cotypes” 4♂♂ (ZMH 61490–ZMH 61493) (Fig. 50). “CALABRIA / bassingliatella oder Cassugliatella / 6.VIII.21 / ? // v. silaecola / Vrty // “Cotypus” / v. / silaecola / Vrty // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61490”; “CALABRIA / bassingliatella oder Cassugliatella / 6.VIII.21 / ? // “Cotypus” / v. / silaecola / Vrty // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61491”; “CALABRIA / bassingliatella oder Cassugliatella / 1.VIII.21 / ? // “Cotypus” / v. / silaecola / Vrty // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61492”; “CALABRIA / Bassigliabllo / 10.VII.21 / ? // “Cotypus” / v. / silaecola / Vrty // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61493”.

Original locality

Italy: Calabria, Cosenza, La Sila [for Anthrocera meliloti silana Turati, 1923]

Current status

Synonym of Z. viciae sicula Calberla, 1895.

Remarks

Zygaena meliloti (Esper, 1789) is a junior synonym to Zygaena viciae (Denis & Schiffermüller, 1775). The taxon silaecola Verity, 1930 was proposed as an objective replacement name for Anthrocera meliloti silana Turati, 1923, which is a junior secondary homonym to Zygaena lonicerae silana Burgeff, 1914. However, the taxon silaecola Verity, 1930 is a junior synonym to sicula Calberla, 1895. The current combination of this taxon is therefore Z. viciae sicula Calberla, 1895 (Hofmann and Tremewan 1996). Moreover, as the reviewer Alberto Zilli correctly states, if the type status of specimen(s) in case an original name has objectively been replaced which is the case here as the name silaecola Verity, 1930 is stated to be a replacement for silana Turati, 1923 due to the latter entering homonymy with another name. In this case, ICZN article 72.7 clearly state that the type specimens remain those of the original name regardless of any statement to the contrary (e.g. other specimens named as types). Accordingly, the listed ‘types’ are such only if they are part of the original Turati’s series, if they are part of another series worked upon by Verity they are not true types of silaecola. Consequently, the ‚’types‘ listed above are incorrectly labelled as types.

Zygaena (Zygaena) meliloti subglockneriana f. charonides Reiss, 1943

Zygaena meliloti ssp. subglockneriana f.charonides Reiss, 1943: Z. Wien. Ent. Ges. 28: 108.

Original material examined

Labelled as “Cotype” 3♂♂ (ZMH 61494–ZMH 61496) (Fig. 51). “CALABRIA / bassingliatella oder Cassugliatella / 4.VIII.21 / ? // f. charonides / Trty // Cotypus / v. silaecola / f. charonides / Trti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61494”; “CALABRIA / Jossiata / 6.VIII.21 / A.FIGRI // Cotypus / v. silaecola / f. charonides / Trti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61495”; “CALABRIA / bassingliatella oder Cassugliatella / 4.VIII.21 / ? // Cotypus / v. silaecola / f. charonides / Trti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61496”.

Original locality

‘Kleine Fleiß, Großglocknergebiet’ [Austria: Carinthia, Grossglockner Region, Kleines Fleisstal].

Current status

Infrasubspecific and hence unavailable name.

Remarks

The taxon charonoides Reiss, 1943 was described as a ‘forma’ (an aberration) of Zygaena meliloti subglockneriana Reiss 1943 (now Z. viciae subglockneriana Reiss 1943 (Hofmann and Tremewan 1996)). According to Article 45.6.4., the author expressly gave it infrasubspecific rank, in which case it is infrasubspecific, and hence unavailable.

Zygaena nevadensis kricheldorffi Reiss, 1933

Zygaena nevadensis kricheldorffi Reiss, 1933: Die Gross-Schm. der Erde, Sup. 2: vii: 252.

Type material examined

Syntypes 2♂♂ (ZMH 61499–ZMH 61500) (Fig. 52). “Cintra / Lusitania / A. Kricheldorff // Cotypus / ssp. kricheldorffi / Reiß // ssp. kricheldorffi / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61499”; “Guarda / Lusitania / A. Kricheldorff // Cotypus / Z. nevadensis / ssp. kricheldorffi / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61500”.

Type locality

Beira Alta, Guarda [Portugal].

Current status

Synonym of Z. nevadensis schmidti Reiss, 1931.

Remarks

The taxon kricheldorffi Reiss, 1933 was described as a subspecies of Z. (Silvicola) nevadensis Rambur, 1858. Hofmann and Tremewan (1996) treated it as a synonym of Z. nevadensis schmidti Reiss, 1931 (type locality: Spain: Avila, Sierra de Gredos, vicinity of Arenas de San Pedro), which is the current valid combination.

Zygaena nevadensis falleriana Reiss, 1931

Zygaena nevadensis falleriana Reiss, 1931: Int. Ent. Z. 25: 111.

Type material examined

Syntype 1♂ (ZMH 61501) (Fig. 53). “falleriana Reifs I / Albarracin / leg. faller 20.8.30 // Cotype m / falleriana / Reifs / H.Reiss // ssp. falleriana / Reifs // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61501”.

Type locality

Sierra Noguera, Albarracin [Spain: Teruel].

Current status

Synonym of Z. nevadensis nevadensis Rambur, 1858.

Remarks

The taxon falleriana Reiss, 1931 is a junior synonym of the nominotypical Z. nevadensis nevadensis Rambur, 1858 (Hofmann and Tremewan 1996).

Zygaena nevadensis falleriana ab. confluens Reiss, 1936

Zygaena nevadensis ssp. falleriana ab. confluens Reiss, 1936: Entomologische Rundschau 54: 29.

Original material examined

Labelled as “Type / Cotype” 2♂♂ (ZMH 835542–ZMH 835543) (Fig. 54). “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type m / falleriana Reiß / ab. confluens / Reiß / 1936 H.Reiss // ZMH 835542”; “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Cotype // ZMH 835543».

Original locality

‘Sierra Alte de Albarracin, 1750 m’ [Spain: Teruel, Albarracin, Sierra Alta].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1936) proposed this name as an aberration of Z. nevadensis falleriana Reiss, 1931. Therefore, as stated by Article 45.6.1 (ICZN 1999) it is deemed to be infrasubspecific name – if its author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena nevadensis falleriana ab. amplomaculata Reiss, 1936

Zygaena nevadensis ssp. falleriana ab. amplomaculata Reiss, 1936: Entomologische Rundschau 54: 29.

Original material examined

Labelled as “Type / Cotype” 4♂♀ (ZMH 835547–ZMH 835550) (Fig. 55). “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type m / falleriana / Reiß ab. amplo- / maculata Reiß / 1936 H.Reiss // ZMH 835547”; “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Cotypen // ZMH 835548»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835549»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type f / falleriana Reiß / ab. amploma- / culata Reiß / 1936 H.Reiss // ZMH 835550»

Original locality

‘Sierra Alte de Albarracin, 1750 m’ [Spain: Teruel, Albarracin, Sierra Alta].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1936) proposed this name as an aberration of Z. nevadensis falleriana Reiss, 1931. Therefore, as stated by Article 45.6.1 (ICZN 1999) it is deemed to be infrasubspecific name – if its author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena nevadensis schmidti Reiss, 1931

Zygaena nevadensis schmidti Reiss, 1931: Int. Ent. Z. 25: 112.

Type material examined

Syntype 1♂ (ZMH 835551) (Fig. 56), “VI. 1927 / Arenas S. Padeo / Prov. Avila // Cotype // Cotype m / schmidti / Reiß / H.Reiss // ZMH 835551”.

Type locality

‘Vorberge der Sierra de Gredos Umgebung Arenas St. Pedro, Prov. Avila, westlich Madrid’ [Spain: Sierra de Gredos, Avila, vicinity of Arenas de San Pedro] (Hofmann and Tremewan 1996).

Current status

Valid subspecies.

Figures 47–56.

47. Zygaena (Agrumenia) truchmena magnifica Reiss 1935; 48. Zygaena (Zygaena) lonicerae stettinensis Reiss, 1922; 49. Zygaena (Zygaena) lavandulae barcelonica Reiss, 1936; 50. Zygaena (Zygaena) meliloti silaecola Verity, 1930; 51. Zygaena (Zygaena) meliloti subglockneriana f. charonides Reiss, 1943; 52. Zygaena nevadensis kricheldorffi Reiss, 1933; 53. Zygaena nevadensis falleriana Reiss, 1931; 54. Zygaena nevadensis falleriana ab. confluens Reiss, 1936; 55. Zygaena nevadensis falleriana ab. amplomaculata Reiss, 1936; 56. Zygaena nevadensis schmidti Reiss, 1931.

Zygaena zapateri Reiss, 1936

Zygaena zapateri Reiss, 1936: Entomologische Rundschau 54: 57.

Type material examined

Syntypes 4♂♂ (ZMH 835552–ZMH 835555) (Fig. 57). «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Cotypen // ZMH 835552»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835553»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835554»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835555”.

Type locality

‘Albarracin’ [Spain: Teruel, vicinity of Albarracin] (Hofmann and Tremewan 1996).

Current status

Synonym of Z. sarpedon carmencita Oberthür, 1910.

Remarks

The taxon Z. zapateri was synonymised with Z. sarpedon carmencita Oberthür, 1910 by Hofmann and Tremewan (1996).

Zygaena sarpedon leuzensis ab. quinquepuncta Reiss, 1958

Zygaena sarpedon leuzensis ab. quinquepuncta Reiss, 1958: Bulletin de la Société entomologique de Mulhouse 1958: 47.

Original material examined

Labelled as “Cotype” 3♂♂ (ZMH 835556–ZMH 835558) (Fig. 58). «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Cotypen // ZMH 835556»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835557”; “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835558”.

Original locality

Not mentioned. The three syntypes are from the „Sierra Alta de Albarracin“.

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1958) proposed this name as an aberration of Z. s. leuzensis Dujardin, 1956. Therefore, as stated by Article 45.6.1 (ICZN 1999) it is deemed to be infrasubspecific name – if its author expressly gave it infrasubspecific rank – which is hence unavailable.

Zygaena zapateri ab. rubrior Reiss, 1958

Zygaena zapateri ab. rubrior Reiss, 1958: Bulletin de la Société entomologique de Mulhouse 1958: 47.

Original material examined

Labelled as “Type / Cotype” 3♂♂ (ZMH 835559–ZMH 835561) (Fig. 59). “Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type m / zapateri ab / rubrior Reiß / 1936 / H. Reiss // ZMH 835559”; “Dr. Geipke // Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Cotypen // ZMH 835560”; “Dr. Geipke // Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835561”.

Original locality

“Sierra Alta de Albarracin” [Spain: Teruel, Albarracin, Sierra Alta, 1700 m].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1958) proposed this name as an aberration of Z. zapateri Reiss, 1936. Therefore, it is deemed to be infrasubspecific, it is hence unavailable (Article 45.6.2, ICZN 1999).

Zygaena achilleae ab. rubrescens Reiss, 1922

Esp. ab. rubrescens Reiss, 1922: Int. Ent. Z. 16: 84.

Original material examined

Labelled as “Type / Cotype” 5♂♂ (ZMH 835564–ZMH 835568) (Fig. 60). «Italia central. / 15/7 Majella 27 / 2400m / coll. F. Dannehl // 15.7.1927 / majella / Central-Italien // ZMH 835563»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type f / aragoneusis / Bgr. ab. rubres- / cens Reiß / 1936 H. Reiss // ZMH 835564»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835565»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Cotypen // ZMH 835566»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835567»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835568».

Original locality

Sierra Alta de Albarracin” [Spain: Teruel, Albarracin, Sierra Alta, 1700 m].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Reiss (1922) proposed this name as an aberration of Zygaena achilleae (Esper, 1780). Therefore, it is deemed to be infrasubspecific, it is hence unavailable (Article 45.6.2, ICZN 1999).

Zygaena trifolii noguerensis Reiss, 1936

Zygaena trifolii var. noguerensis Reiss, 1936: Entomologische Rundschau 54: 89.

Type material examined

Syntypes 4♂♀ (ZMH 835569–ZMH 835572) (Fig. 61). «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type m / noguerensis / Reiß / 1936 H. Reiss // ZMH 835569»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Type // Type f / noguerensis / Reiß / 1936 H. Reiss // ZMH 835570»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // Cotypen // ZMH 835571»; «Sierra Alta de / Albarracin 1700 m / 5–25.7.35. / coll. Herbert Noack. // ZMH 835572».

Type locality

‘Sierra Alta de Albarracin, 1750 m’ [Spain: Teruel, Sierra Alta, Albarracín] (Hofmann and Tremewan 1996)

Current status

Synonym of Zygaena trifolii barcelonensis Reiss, 1922.

Remarks

Reiss (1936) proposed this name as a variation of Z. trifolii (Esper, 1783) from Teruel, Sierra Alta, Albarracín (Spain). According to Article 45.6.4., it is subspecific if first published before 1961 and author expressly used the terms “variety”. The taxon noguerensis Reiss, 1936 is a junior synonym to Zygaena trifolii barcelonensis Reiss, 1922 (Hofmann and Tremewan 1996).

Zygaena (Agrumenia) occitanica tourettica Reiss, 1953

Zygaena occitanica tourettica Reiss, 1953: Entomologische Zeitschrift. Frankfurt a. M. 63: 111.

Type material examined

Syntypes 1♂ 1♀ (ZMH 61533–ZMH 61534) (Fig. 62). “22306. // occitanica-tourrettica // Cotype m / tourettica / Reiß / 1953 H. Reiß //Alp. Maritimes / Tourrettessur / Loiys / 27.6.1951 / H. Reiß Stuttgart // Coll. K. Albrecht / Eing. Nr. 2, 57 // Alpes-Maritimes / Tourette 360m, 27.6.51// Alpes-Maritimes / Touvette 360m, 27.6. / 1951 // ZMH 61533”; “O.lp. Maritimes / Tourrettess. Loup / 360m 1.7.1951 / H. Reiß Stuttgart // 22307. // Cotype f / tourettica / Reifs / 1953 H.Reiß // Alpes-Miritimes / Torette Tourette 27.6.51 // Coll. K. Albrecht / Eing. Nr. 2, 57 // ZMH 61534”.

Type locality

Tourettes-sur-Loup [France: Alpes-Maritimes].

Current status

Synonym of Z. occitanica praematura Przegendza, 1932.

Remarks

The taxon tourettica Reiss, 1953 is a junior synonym to Z. occitanica praematura Przegendza, 1932 (Hofmann and Tremewan 1996).

Zygaena (Agrumenia) transpamirina nuksanensis Koch, 1937

Zygaena nuksanensis Koch, 1937: Entomologische Zeitschrift. Frankfurt a. M. 51: 61.

Type material examined

Syntype 1♂ (ZMH 61535) (Fig. 63). “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500-4000m Mittel Juli / leg. H. & E. Kotzsch // Co-Type / e Collection / Manfred Koch // nuksanensis / Koch // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61535”.

Type locality

Noqşan An [Afghanistan: Badakhshan, NE Hindukush, Nuksan-Pass].

Current status

Valid subspecies.

Remarks

Zygaena nuksanensis Koch, 1937 is currently considered to represent a valid subspecies of Z. transpamirina Koch, 1936 (Hofmann and Tremewan 1996).

Zygaena (Mesembrynus) punctum kolbi Reiss, 1933

Zygaena (Hyala) punctum var. kolbi Reiss, 1933: Die Gross-Schm. der Erde, Sup. 2: vii: 255.

Type material examined

Syntypes 2♂♂ (ZMH 61545–ZMH 61546) (Fig. 64). „Dalmatien / Susak Trsat / 25.VI.31. / 140m No1708 / Kolb, München // CO-TYPE VON Zyg. punctum / v. Kolbi Reiss. / DANIEL / MÜNCHEN // var. Kolbi / Reifs // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61545“; „Dalmatien / Susak Trsat / 25.VI.31. / 140m No1708 / Kolb, München // CO-TYPE VON Zyg. punctum / v. kolbi Reiss. / DANIEL / MÜNCHEN // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61546”.

Type locality

‘Fiume (Susak Trsat)’ [Croatia: vicinity of Rijeka, Susak Trsat].

Current status

Synonym of Z. punctum itala Burgeff, 1926.

Remarks

Reiss (1933) proposed this name as a variation of Z. punctum Ochsenheimer, 1808 from Fiume (Croatia). According to Article 45.6.4., it is subspecific if first published before 1961 and author expressly used the terms “variety”. The taxon kolbi Reiss, 1933 was synonymised with Z. punctum itala Burgeff, 1926 by Hofmann and Tremewan (1996). This is the current valid combination. However, the taxonomic position of the subspecies of Z. punctum from the Balkans is currently again revised (Nahirnić, Jakšić and Tarmann, in preparation).

Zygaena (Zygaena) persephone Zerny, 1934

Zygaena persephone Zerny, 1934: Z. Öst. Ent. Ver. 19: 108, 200.

Type material examined

Paratype 1♂ (ZMH 61547) (Fig. 65). “H.Atlas Maroc. / Tizi u Tachdirt / 3000–3200 m VII.33 / Schwingenschuss // Zygaena / persephone / Zerny / Paratype // persephone / Zerny // Coll Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61547”.

Type locality

‘Großer Atlas, Tizi’n Tachedirt (auch Tizi’n Yabessen genannt)’ [Morocco: Haut-Atlas, Djebel Toubkal region, Tizi-n-Tacheddirt].

Current status

Valid species.

Zygaena (Zygaena) transalpina asturiensis Reiss, 1936

Zygaena transalpina asturiensis Reiss, 1936: Entomologische Rundschau 54: 91.

Type material examined

Syntypes 3♂♂ (ZMH 61548 – ZMH 61550) (Fig. 66). “Cotypus / transalpina ssp. asturiensis / Reifs // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61549”; “Picos de Europa / La Liebana / A. Kricheldorff // Cotypus / transalpina ssp. asturiensis / Reifs // ssp. asturiensis / Reiß // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61548”; “Picos de Europa / La Liebana / A. Kricheldorff // Cotypus / transalpina ssp. asturiensis / Reifs // Coll Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61550”.

Type locality

‘Picos de Europa in Nordspanien, La Liebana’ [Spain: Picos de Europa, Cantabria, vicinity of la Vega de Liébana] (Hofmann and Tremewan, 1996).

Current status

Synonym of Z. transalpina rupicola Rocci, 1936.

Remarks

A junor synonym to Z. transalpina rupicola Rocci, 1936. Rocci’s work was pulished in May 1936, whereas the paper of Reiss (1936) in November 1936 (Hofmann and Tremewan 1996).

Figures 57–66.

57. Zygaena zapateri Reiss, 1936; 58. Zygaena sarpedon leuzensis ab. quinquepuncta Reiss, 1958; 59. Zygaena zapateri ab. rubrior Reiss, 1958; 60. Zygaena achilleae ab. rubrescens Reiss, 1922; 61. Zygaena trifolii noguerensis Reiss, 1936; 62. Zygaena (Agrumenia) occitanica tourettica Reiss, 1953; 63. Zygaena (Agrumenia) transpamirina nuksanensis Koch, 1937; 64. Zygaena (Mesembrynus) punctum kolbi Reiss, 1933; 65. Zygaena (Zygaena) persephone Zerny, 1934; 66. Zygaena (Zygaena) transalpina astu riensis Reiss, 1936.

Zygaena (Agrumenia) sogdiana altissima Burgeff, 1914

Zygaena fraxini var. altissima Burgeff, 1914: 51.

Type material examined

Syntypes 8♂♀ (ZMH 61552–ZMH 61559) (Fig. 67). “Alai mont. / 1905 Korb // Cotypus / altissima / Bgff // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61552”; “Alai mont. // Cotypus / sogdiana / altissima / Bgff // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61553”; “Alai mont. / 1905 Korb // Cotypus / sogdiana / altissima / Bgff // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61554”; “Alai mont. / 1905 Korb // Cotypus / sogdiana / altissima / Bgff // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61555”; “Alai mont. / 1905 Korb // Cotype f / altissima / Bgff / H.Reiss // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61556”; “Alai mont. // Cotype m / altissima / Bgff / H. Reiss // v. altissima / Bgff // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61557”; “Alai mont. / 1905 Korb // Cotypus / altissima / Bgff // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61558”; “Alai mont. / 1905 Korb // Cotypus / sogdiana / altissima / Bgff // Coll Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61559”.

Type locality

‘Trans-Alaikette, Ak-Bassegha, in einer Höhe von 2000 m’ [TransAlai near Tajikistan border].

Current status

Valid subspecies.

Remarks

Burgeff (1914) proposed this name as a variation of Z. fraxini Ménétriés, 1832 from Tajikistan. According to Article 45.6.4., it is subspecific if first published before 1961 and author expressly used the terms “variety”. The taxon altissima Burgeff, 1914 has been transferred to Zygaena ransalp Erschoff, 1874. The current combination is Z. ransalp altissima Burgeff, 1914 (Hofmann and Tremewan 1996).

Zygaena (Agrumenia) rosinae Korb, 1903

Zygaena rosinae Korb, 1903: 326.

Type material examined

Syntypes 2♂♂ (ZMH 61569–ZMH 61570) (Fig. 68). “R. drenonien / Kulp. / Korb // Rosinae // Cotype m / rosinae / Korb // H.Reiss. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61569”; “R. Armen / Kulp. / 1901 Korb // Cotypus / Z. rosinae / m Korb // rosinae / Korb. // Coll. Bytinski-Salz / Eing Nr. 20, 1960 // ZMH 61570”.

Type locality

‘Russisch-Armenien, Kulp’ [Turkey: Kars, Tuzluca].

Current status

Valid species.

Zygaena (Zygaena) transalpina sorrentina f. aureomaculata Stauder, 1921

Zygaena transalpina sorrentina forma aureomaculata Stauder, 1921: 30

Original material examined

Labelled as “Type” 1♂ (ZMH 61573) (Fig. 69). “Type // 1/6 / Ital. mer. Goo m. / penins. Surrentina / M. Faito 1921 oder / / H. Stauder legit. // f. aureomacula / ta Hauder // aureomaculata // Sig. G. Warnecke / Eing Nr 5, 1949 // ZMH 61573”.

Original locality

’M. Faito bei Castellamare di Stabia, 800–900 m’ [Italy: Campania, Napoli, Monte Faito, south of Castellamare di Stabia].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Described as a golden-yellow form of Zygaena ransalpine sorrentina with a remark by Stauder (1921) that this form may be an aberration caused by high temperature. According to Article 45.6.4., the author expressly gave it infrasubspecific rank, in which case it is infrasubspecific, and hence unavailable.

Unpublished, unavailable names

Zygaena exulans v. amarensis Diehl

Material examined

Labelled as “Cotypes” 2♂♂ (ZMH 835562–ZMH 835563) (Fig. 70). “Italia central. / 27 Majella 15/7 / 2400m / coll. F. Dannehl // Cotypen // 15.7.1927 / majella / Central-Italien // ZMH 835562”; “Italia central. / 15/7 Majella 27 / 2400m / coll. F. Dannehl // 15.7.1927 / majella / Central-Italien // ZMH 835563”.

Current status

“in litteris” name and hence not available.

Remarks

This taxon was never published. Diehl originally intended to describe the population from the Abruzzi in central Italy. Later he obviously realised that Burgeff (1926) had already named it a year before Dannehl collected these two specimens (1927). But even Burgeff’s name (subsp. Abruzzina) is not based on the discovery, as it was (and still is) a ransalpin name of a former homonymic name (subsp. Apennina Rebel, 1910, preoccupied by apennina Turati, 1884, a subsp. Of Z. carniolica) (Axel Hofmann pers. comm.).

Zygaena (Zygaena) transalpina nigraltudinaria ab. Flava Rocci, 1937

Material examined

Labelled as “Type” 1♂ (ZMH 61551) (Fig. 71). “Breth / Küstenland / 9/8 05 // ex. Coll. / Ronnicke / TYPE // ab. Flava. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61551”.

Current status

“in litteris” name and hence not available.

Remarks

There is no Zygaena r ansalpine hilfi ab. Flava Rocci, 1936 published anywhere. Therefore, this material is no type-material in the sense of the Zoological Nomenclature.

Zygaena (Zygaena) transalpina sorrentina v. eranesceus Sicher, 1906

Material examined

Labelled as “Type” 1♂ (ZMH 61572) (Fig. 72). “Type // 1/6 / Ital. mer. Goo m. / penins. Sorrento / M.Faito 192 1 oder / / H. Stauder legit. // f. eranesceus / Stauder // Sig. G Warnecke / Eing Nr 5, 1949 // // ZMH 61572“.

Current status

“in litteris” name and hence not available.

Remarks

This taxon is missing in all know catalogues. In Sicher (1906) it is not mentioned either. Mag. H. Bruckner from the library of the Natural History Museum in Vienna (Naturhistorisches Muse7m, Wien, Austria) checked every page of the volume of the Jahresberichte des Wiener Entomologischen Vereins, Volume 16 (1905) and could not find any other paper, nor note from Sicher. We think that the specimen was pre-labelled as type, but Sicher finally did not describe it.

Zygaena (Zygaena) transalpina litorea v. flava Stauder

Material examined

Labelled as “Cotypes” 5♂♀ (ZMH 61574–ZMH 61578) (Fig. 73). « ex. flava / e.l. / 11.4.30. // ex. coll. Ronnicke / Cotype // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61574 » ; « Italia / Brescia 19 / P. Ronnicke // ex. flava / e.l. / 30.6.30. // ex coll. / Ronnicke / Cotype // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61575 » ; « Italia / Brescia 19 / P. Ronnicke // ex. flava / e.l. / 5.6.30. // ex coll. / Ronnicke / Cotype // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61576 » ; « ex. flava / e.l. / 11.6.30. // ex. coll. Ronnicke / Cotype // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61577 » ; « Italia / Brescia 19 / P. Ronnicke // ex. flava / e.l. / 4.6.30. // ex coll. / Ronnicke / Cotype // ab. Flava / Ronnicke // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61578”.

Current status

“in litteris” name and hence not available.

Remarks

In all pubilcations of Stauder there is no such taxon mentioned.

Zygaena (Mesembrynus) purpuralis v. asupinensis Bytinski-Salz

Material examined

Labelled as “Cotypes” 9♂♀ (ZMH 61536–ZMH 61544) (Fig. 74). “Rovigno d´lstria / 6.1934 / Dr. H. Bytinski-Salz // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61536”; “Rovigno d´lstria / 6.1934 / Dr.H.Bytinski-Salz // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61537”; “Rovigno d´lstria / 6.1934 / Dr. H. Bytinski-Salz // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61538”; “Rovigno d´lstria / 6.1934 / Dr.H.Bytinski-Salz // Cotypus // var. asupinensis / By.5. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61539”; “Rovigno d´lstria / 6.1933 / Dr. H. Bytinski-Salz // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61540”; “Rovigno d´lstria / 6.1934 / Dr. H. Bytinski-Salz // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61541”; “Rovigno d´lstria / 1.–6.6.1933 / Dr. H. Bytinski-Salz // Cotypus // Coll. Bytinski-SalzColl. / Eing. Nr. 20, 1960 Eing. // ZMH 61542”; “Brioni / Istrien / 28.V.1934 / Dr. H. Bytinski-Salz // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61543”; “Rovigno d´lstria / 1.–6.VI.1933 / Dr. H. Bytinski-Salz // Cotypus // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61544”.

Current status

“in litteris” name and hence not available.

Remarks

This taxon cannot be found in the literature.

Figures 67–74.

67. Zygaena (Agrumenia) sogdiana altissima Burgeff, 1914; 68. Zygaena (Agrumenia) rosinae Korb, 1903; 69. Zygaena (Zygaena) transalpina sorrentina v. aureomaculata Stauder, 1921; 70. Zygaena exulans v. amarensis Diehl; 71. Zygaena (Zygaena) transalpina nigraltudinaria ab. flava Rocci, 1937; 72. Zygaena (Zygaena) transalpina sorrentina v. eranesceus Sicher, 1906; 73. Zygaena (Zygaena) transalpina litorea v. flava Stauder; 74. Zygaena (Mesembrynus) purpuralis v. asupinensis Byt.-S.

Subfamily Chalcosiinae Walker, 1865

Aglaope labasi meridionalis Zerny, 1934

Aglaope labasi meridionalis Zerny, 1934: Z. Öst. Ent. Ver. 19: 29

Type material examined

Paratype 1♂ (ZMH 61474) (Fig. 75). “H. Atlas Maroc. / Tachdirt 2300– / 2700 m 3.–25.VII.33 / Schwingenschuss // Aglaope / labasi / ssp. meridio- / nalis Zerny / PARATYPE // ssp. meridionalis / Zerny // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61474”.

Type localitiy

‘Auf dem Wege nach Tachdirt zum Tiz’n Tachdirt’ [Morocco: Haut-Atlas, Djebel Toubkal region, between Tacheddirt and Tizi-n-Tacheddirt, 2700–2800 m].

Current status

Valid subspecies.

Subfamily Procridinae Boisduval, 1828

Adscita alpina (Alberti, 1937)

Procris alpina Alberti, 1937: Ent. Z. 50 (1936): 435

Type material examined

Paratype 1♂ (ZMH 61373) (Fig. 76). “Paratypus // ? / 28.7.? // Sammlung / Gustav Meyer // Procris / alpina Alb. / m / det.B.Alberti // ZMH 61373”

Type locality

‘Südtirol, Stifser-Joch-Straße, Franzenshöhe, 2000 m [Italy, province Bozen/Bolzano].

Current status

Valid species.

Adscita statices drenowskii (Alberti, 1939)

Procris drenowskii Alberti, 1939: Izv. Tsarsk. Prirod. Inst. Sofia 12: 43.

Type material examined

Paratypes 3♂♂ (ZHM 61397–ZMH 61399) (Fig. 77). “Asia min.c. / Ak-Chehir / 1.–15.Mai / Coll. Wagner-Wien // Procris m / drenowskii / det. B.Alberti // Paratype / Akschekir // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61397”; “Asia min.c. / Ak-Chehir / 1.–15.Mai / Coll. Wagner-Wien // Procris m / drenowskii / det. B. Alberti // Procris m / drenowskii / det. B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61398”; “Asia min. c. / Ak-Chehir / 1.–10.Juni 28 / Coll. Wagner-Wien // f Procris / abscirra Z. ? / det. A. Naufock / 4. 1. 1929 // Procris m / drenowskii / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61399”.

Type locality

Greece: Chalkidiki, Athos.

Current status

Valid subspecies.

Remarks

Alberti (1939) proposed this name as a new species of Procris. This taxon was subsequently downgraded to subspecific rank (Daniel, Forster and Osthelder 1951).

Jordanita graeca persica (Alberti, 1938)

Procris graeca persica Alberti, 1938: Mit. der Mün. Ent. G. 27: 125.

Type material examined

Holotype 1♂ (ZMH 61408) (Fig. 78) Allotype 1♀ (ZMH 61409) Paratypes 2♂♂ (ZMH 61410–ZMH 61411). “Keredi / 1800 256b // IRAN / Keredj (1200m.) / 1936 / Brandt // 303 // ssp. persica / Alberti // Senikl-Präp- / Procris No / By. 3 // Holotype / m / Persien / Keredj // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61408”; “IRAN / Keredj (1200m.) / 1936 / Brandt // 1800 / 146 // Procris gracia / f / det. B. Alberti // Allotype / Persien / Keredj // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61409”; “Iran // Procris gracia / ssp. persica / B. Alberti / 1937 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61410”; “Iran // Paratype / m / Persien / Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61411”.

Type locality

Iran, Keredj [Karaj, Alborz province].

Current status

Valid subspecies.

Remarks

Alberti (1938a) proposed this name as subspecies of P. graeca Jordan, 1907, that is considered a valid subspecies name.

Adscita geryon orientalis (Alberti, 1938)

Procris geryon orientalis Alberti, 1938: Z. für Natur., Halle a. S. 92: 54.

Type material examined

Paratypes 25♂♀ (ZMH 61418–ZMH 61442) (Fig. 79). “Bosnia / Korićna / coll. O. Leonhard // no 46 m / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp. / orientalis / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61418”; “Bosnia 1904 / Korićna / 8.7 / O. Leonhard // no 50 f / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris geryon / ssp. orientalis / B. Alberti 1938 f // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61419”; “Bosnia / Maklen Pass / coll. O. Leonhard // no 70 m / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp / orientalis / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61420”; “Bosnia / Maklen Pass / coll. O. Leonhard // no 64 m / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp / orientalis / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61421”; “Bosnia / Maklen Pass / coll.O.Leonhard // no 77 m / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp / orientalis / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61422”; “Bosnia / Maklen Pass / coll. O. Leonhard // no 69 m / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp / orientalis / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61423”; “Velebit Ostaria / 6.7. / M.Hilf 1910 / Coll. O. Leonhard // no 81 m / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp. / orientalis / B.Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61424”; “Velebit Ostaria / 6.7. / M.Hilf 1910 / Coll. O. Leonhard // no 82 f / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris geryon / ssp. orientalis f / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61425”; “Bosnia 1904 / Korićna / 10.7 / O. Leonhard // no 51 f / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris geryon / ssp. orientalis / B. Alberti f / 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61426”; “Bosnia / Korićna / coll. O. Leonhard // no 43 m / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp. / orientalis / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61427”; “Bosnia / Maklen Pass / coll. O. Leonhard // no 60 f / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris geryon / ssp. orientalis f / B.Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61428”; “Bosnia 1904 / Korićna / 8.7 / O. Leonhard // no 49 m / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris geryon ssp. / orientalis / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61429”; “Bosnia / Maklen Pass / coll. O. Leonhard // no 71 m / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp / orientalis / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61430”; “Bosnia / Korićna / coll. O. Leonhard // no 55 f / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris geryon / ssp. orientalis f / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61431”; “Bosnia 1904 / Korićna / 10.7 / O. Leonhard // no 52 f / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris geryon / ssp. orientalis f / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61432”; “Velebit Ostaria / 18.6 / M.Hilf 1910 / Coll. O. Leonhard // no 80 m / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp. / orientalis / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61433”; “Bosnia / Korićna / coll. O. Leonhard // no 54 f / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris geryon / ssp. orientalis / B. Alberti 1938 f // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61434”; “ ? // Bosnia / Korićna / coll. O. Leonhard // no 59 f / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris geryon / ssp. orientalis f / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61435” : “Bosnia / Maklen Pass / coll. O. Leonhard // no 68 m / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp / orientalis / B.Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61436”; “Bosnia / Korićna / coll. O. Leonhard // no 44 m / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp. / orientalis / B. Alberti 1938 // ssp. orientalis / Alb. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61437”; “Bosnia / Korićna / coll.O.Leonhard // no 58 f / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp. / orientalis / B. Alberti 1938 // ssp. orientalis / Alb. // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61438”; “Bosnia / Korićna / 10.7 / O.Leonhard // no 53 f / Pr. geryon Hb. / det. A.Naufock / VIII 1935 // Procris geryon / ssp. orientalis / B. Alberti 1938 f // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61439”; “Bosnia / Korićna / coll. O. Leonhard // no 57 f / Pr. geryon Hb. / det. A.Naufock / VIII 1935 // Procris geryon / ssp. orientalis f / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61440”; “Bosnia / Korićna / coll. O. Leonhard // no 45 m / Pr. geryon Hb. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp. / orientalis / B. Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61441”; “Bosnia / Maklen Pass / coll. O. Leonhard // no 65 m / Pr. geryon HG. / det. A. Naufock / VIII 1935 // Procris m / geryon ssp / orientalis / B.Alberti 1938 // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61442”.

Type locality

‘Maklenpaß’ [Bosnia and Herzegovina: Maklen Pass].

Current status

Valid subspecies.

Remarks

Alberti (1938b) proposed this name as a subspecies of A. geryon (Hübner, 1813), that is considered a valid subspecies name.

Jordanita (Jordanita) globulariae bosniaca (Alberti, 1937)

Procris globulariae bosniaca Alberti, 1937: Int. Ent. Z. 51: 99

Type material examined

Paratypes 17♂♀ (ZMH 61443–ZMH 61459) (Fig. 80). “PARATYPE m / Procris globulariae / ssp. bosniaca // ? // Sammlung / Gustav Meyer // ZMH 61443”; “Velebit Ostaria / 6.7. / M. Hilf 1910 / Coll. O. Leonhard // no 84 Procrie / cognota K.S. m / det. A. Naufock / 1935. VIII. // ssp. bosniaca / Alberti // Prorris globulariae / ssp bosniaca m / B.Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61444”; “Bosnia 1904 / Korićna 10.7 / O. Leonhard // no 19 f / Pr. cognata HS. / det. A. Naufock / VIII.1935 // Procris globula- / riae ssp. / bosniaca f / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61445”; “Bosnia / Korićna / coll.O.Leonhard // no 8 f / Pr. cognata H.S. / det. A.Naufock / VIII 1935 // Prorris globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61446”; “Bosnia 1904 / Korićna 28.7 / O.Leonhard // no 20 f / Pr. cognata HS. / det. A.Naufock / VIII.1935 // Prossis globu- / lariae ssp. / bosniaca f / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61447”; “Bosnia / Korićna / coll.O.Leonhard // no 6 m / Proar. cognata HS. / det. A.Naufock / VIII.1935 // Prossis globula- / riae ssp. / bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61448”; “Bosnia / Korićna / coll.O.Leonhard // no 10 m / Pr. cognata HS. / det. A.Naufock / VIII.1935 // Prossis globulari- / ae ssp. bosniaca / B. Alberti m // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61449”; “Bosnia / Korićna / coll.O.Leonhard // no 11 m / Pr. cognata HS. / det. A.Naufock / VIII.1935 // Prossis globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61450”; “Bosnia / Korićna / coll.O.Leonhard // no 16 / Pr. cognata HS. / det. A.Naufock / VIII.1935 // Prossis globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61451”; “Bosnia / Korićna / coll.O.Leonhard // no 13 m / Pr. cognata HS. / det. A.Naufock / VIII.1935 // Prossis globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61452”; Bosnia / Korićna / O.Leonhard // no 17 m / Pr. cognata HS. / det. A.Naufock / VIII.1935 // Prossis globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61453”; “Bosnia / Korićna / coll.O.Leonhard // no 14 / Pr. cognata HS. / det. A.Naufock / VIII.1935 // Prossis globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61454”; “Bosnia / Korićna / coll.O.Leonhard // no 7 m / Pr. cognata HS. / det. A.Naufock / VIII.1935 // Prossis globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61455”; “Bosnia / Korićna / coll. O. Leonhard // no 12 m / Pr. cognata HS. / det. A. Naufock / VIII.1935 // Procris globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61456”; “Bosnia / Korićna / coll. O. Leonhard // no 15 m / Pr. cognata HS. / det. A. Naufock / VIII.1935 // Procris globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61457”;”Bosnia 1904 / Korićna 10.7 / O. Leonhard // no 18 m / Pr. cognata HS. / det. A. Naufock / VIII.1935 // Procris globulari- / ae ssp. bosniaca / B. Alberti m // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61458”; “Bosnia / Korićna / coll. O. Leonhard // no 9 m / Pr. cognata HS. / det. A. Naufock / VIII.1935 // Procris globulariae / ssp. bosniaca m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61459”.

Type locality

‘Korična’ [Bosnia and Herzegovina: Korična].

Current status

Invalid subspecific name. Synonym of Jordanita (Jordanita) globulariae globulariae (Hübner, 1793).

Remarks

Alberti (1937c) proposed this name as subspecies of Procris globulariae Hübner (1793). The name was subsequently synonymized. The current combination is Jordanita (Jordanita) globulariae globulariae (Hübner, 1793) (Efetov and Tarmann 1995).

Procris mauretanica Naufock, 1932

Procris mauretanica Naufock, 1932: Z. Öst. Ent. Ver. 17: 77

Type material examined

Paratypes 2♂♂ (ZMH 61497–ZMH 61498) (Fig. 81). “Mauretania central. / Atlas major / Areg-Ikis / Iminem-Tal.2000 m. / 15.–20.7.1932. / H. Gürck leg. // Pr. mauretanica / m. / det. A. Naufock / 1.II.1935 m // ZMH 61497 // Procris / mauretanica / Naufock / PARATYPE // mauretanica / Nauf // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // 11 / 49”; “Mauretania central. / Atlas major / Areg-Ikis / Iminem-Tal.2000 m. / 15.–20.7.1932. / H. Gürck leg. // mauretanica / ? // Procris / mauretanica / Naufock / PARATYPE // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // 12 / 52 // ZMH 61498”.

Type locality

‘Französisch Marokko, Aiin Leuh, Sefrou’ [Morokko: Moyen Atlas, Ain Leuh and Sefrou].

Current status

Valid species.

Procris statices extensa Alberti, 1937

Procris statices extensa Alberti, 1937c: Int. Ent. Z. 51: 100

Type material examined

Paratypes 9♂♀ (ZMH 61560–ZMH 61568) (Fig. 82). “Transsylvania / ex coll. / Neuburger // Pr. statices / no 35 / L. m / det. Nfk.19VIII5 // 104 // Procris statices / ssp extensa f / B. Alberti ? // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61560”; “mwv? / 117 a // Bosnia / Maklen Pass / coll. O. Leonhard // Pr. statices / no 30 / L. m / det.Nfk.1935 VIII // Procris statices / ssp. extensa m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61561“; „Kuriém / 46609 // Italien // Bosnia / Korićna / coll. O. Leonhard // Pr. statices / no 33 / L. m / det.Nfk.1935 VIII // Procris statices / ssp. extensa m / B. Alberti 1938 // ssp. extensa / Alberti / ZMH 61563”; “mwv? / 1017 w // Pr. statices / no 29 / L. m / det. Nfk.1935 VIII // Bosnia / Maklen Pass / coll. O. Leonhard // Procris statices ssp. extensa / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61564”; “Italien // Bosnia / Maklen Pass / coll. O. Leonhard // Pr. statices / no 26 / L. m / det. Nfk.1935 VIII // Procris statices / ssp. extensa m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61565”; “? // Bosnia / Korićna / coll. O. Leonhard // Pr. statices / no 34 / L. f VIII / det.Nfk.1935 // Procris statices ssp. / extensa f / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61566”; “? //Bosnia / Maklen Pass / coll. O. Leonhard // Pr. statices / no 28 / L. m VIII / det.Nfk.1935 // Procris statices / ssp. extensa m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61567”; “Jno Sda – / Lias Z / masu 27/6.Tz // Pr. statices / no 28 / L. m VIII / det.Nfk.1935 // Procris statices / ssp. extensa m / B. Alberti // Coll. Bytinski-Salz / Eing. Nr. 20, 1960 // ZMH 61568”.

Type locality

‘Maklenpaß’ [Bosnia and Herzegovina: Maklen Pass].

Current status

Invalid subspecies name. Synonym of Adscita (Adscita) statices statices (Linnaeus, 1758).

Remarks

This taxon was subsequently synonymized. The current combination is Adscita (Adscita) statices statices (Linnaeus, 1758) (Efetov and Tarmann 1995).

Theresimima ampellophaga ab . astrapta Dannehl, 1933

Ino ampellophaga ‘ab. (?) rasse (?) astrapta Dannehl, 1933: Entomologische Zeitschrift. Frankfurt a. M. 47: 147

Original material examined

Labelled as “Cotype” 1♂ (ZMH 835573) (Fig. 83). “57 // Italia central. / Mte. Sirente / coll. F. Dannehl // astrapta / Col. // 5.7.1928 / mte sirente / Mittel-Italien // ZMH 835573».

Original locality

‘Sabiner Berge, c. 500 m’ [Italy: Latio, Rieti, Monte Sabini].

Current status

Infrasubspecific and hence unavailable name.

Remarks

Dannehl (1933) proposed this name as an aberration of Theresimima ampellophaga Bayle-Barelle, 1808. Therefore, it is deemed to be infrasubspecific, it is hence unavailable (Article 45.6.2, ICZN 1999).

Figures 75–83.

75. Aglaope labasi meridionalis Zerny, 1934; 76. Adscita alpina (Alberti, 1937); 77. Adscita statices drenowskii (Alberti, 1939); 78. Jordanita graeca persica (Alberti, 1938); 79. Adscita geryon orientalis (Alberti, 1938); 80. Jordanita (Jordanita) globulariae bosniaca (Alberti, 1937); 81. Procris mauretanica Naufock, 1932; 82. Procris statices extensa Alberti, 1937; 83. Theresimima ampellophaga ab. astrapta Dannehl, 1933.

Discussion

More than 2,000 type specimens of butterflies and moths (Lepidoptera) are deposited in the Zoological Museum of Hamburg (ZMH). In this catalogue, we have listed 259 “type” specimens: 13 valid species names, 34 subspecies (16 valid and 18 invalid names), and 36 invalid infrasubspecific rank under the ICZN code. Of these 259 specimens, 81 are primary types, 92 are secondary types, and 86 are considered invalid infrasubspecific rank (Suppl. material 1: Table S1).

Comparison of the first part of our updated catalogue to the type catalogue of Weidner (1974) reveals 32 new additions to the families Sesiidae, Brachodidae, Metarbelidae, and Zygaenidae (Suppl. material 1: Table S1). New additions to the type collection since the Weidner catalogue (1974) do not necessarily reflect “new type additions” – except for the five taxa donated by Kallies and Bartsch (Sesiidae and Brachodidae), and Lehmann (Metarbelidae) to the ZMH collection– they were found among the exisiting type material and were either mididentified or misplaced under the incorrect type labels. Details of the changes and updates including the new types, changes to the type status, and changes to the specimen counts since the last catalogue of Weidner (1974) were summarized in Suppl. material 1: Table S1.

The collection of Dr. Hanan Bytinski-Salz was the largest collection containing the most “type specimens” mainly from Hugo Reiss (89 specimens), Burchard Alberti (59 specimens) and Hanan Bytinski-Salz (35 specimens). None of the specimens from the latter author represents a valid name. The oldest type specimen was from the collection of Bytinski-Salz described by Korb in 1903 (Zygaena rosinae Korb, 1903).

Acknowledgements

This project was funded by the CeNak “Digitalisierung der CeNak-Sammlungen”. We also like to thank HASPA Lotterie Sparen for funding boxes and the City of Hamburg for further funding via Bussgeldfonds. We thank Dr. Henrik Kusche (Leitung Sammlungsmanagement/Digitalisierung) and the Gesellschaft der Freunde und Förderer des Zoologischen Museums Hamburg for providing financial support to the collections. For help with the search of rare literature we thank Mag. Harald Bruckner and Dr. Sabine Gaal (both Naturhistorisches Museum Wien, Austria), Mag. Ursula Grimm (Tiroler Landesmuseen, Ferdinandeum, Innsbruck, Austria) and Prof. Dr Predrag Jakšić (Belgrade, Serbia). We are extremely grateful to Axel Hofmann (ABL – Arten Biotope Landschaft – Bürogemeinschaft für Landschaftsökologie, Freiburg, Germany) for his constructive and fruitful comments with the difficult taxa (unpublished and unavailable names). The authors also acknowledge the members of the other Zoological Collections at the CeNak for their patience during our intensive photography sessions. Comments by Alberto Zilli, Wolfgang Naesig, Roman Yakovlev, and Karen McLachlan Hamilton substantially improved the manuscript.

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