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An illustrated catalogue of the type specimens of Lepidoptera housed in the Zoological Museum Hamburg (ZMH): Part II. superfamily Papilionoidea
expand article infoReza Zahiri§, Vazrick Nazari|, Hossein Rajaei, Martin Wiemers#, Maryam Fatahi, Matthias Seidel, Thure Dalsgaard, Martin Husemann
‡ Leibniz Institute for the Analysis of Biodiversity Change, Hamburg, Germany
§ Canadian Food Inspection Agency, Ottawa, Canada
| Unaffiliated, Ottawa, Canada
¶ State Museum of Natural History Stuttgart, Stuttgart, Germany
# Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany
Open Access

Abstract

We provide an updated catalogue of the type material of the lepidopteran superfamily Papilionoidea deposited in the Zoological Museum of Hamburg (ZMH). We report 414 specimens labelled as “types” belonging to nine species (all of valid names), 74 subspecies (44 valid names and 30 synonyms), 59 invalid infrasubspecific names under the ICZN code, and 23 specimens of 16 “in litteris” (= unavailable) names. Out of the 414 specimens labelled as “types”, 171 specimens are primary types (8 holotypes/lectotypes and 163 syntypes) and 80 are secondary types; 120 specimens are infrasubspecific and hence invalid and are considered as “original specimens”; and 43 specimens are treated here as “Non-type” specimens (topotype). We present a full bibliography of the original descriptions and illustrations for all of these taxa, aiming to provide a comprehensive taxonomic guide to this collection.

Key Words

Biodiversity, CeNak, Herbert Weidner, ZMH, digitization, Georg Warnecke, Hanan Bytinski-Salz

Introduction

Natural history collections are the main archives of biodiversity. They focus on collecting, maintaining and documenting natural specimens, a crucial task, especially in times of global biodiversity decline (Dirzo et al. 2014). The Zoological Museum of Hamburg (ZMH) holds large collections across all animal groups. With approximately five million specimens and several thousand primary types, the entomological collection is among the most important natural history archives in Germany. Yet, the collections are overall not well documented and updated type catalogues are largely lacking.

The last ZMH Lepidoptera catalogue was published by Herbert Weidner, a former curator of the collection, in 1974 (Weidner 1974). Since the publication of this work more than 40 years ago, no further types have been documented. We initiated a comprehensive effort to document all of the type material preserved in the ZMH in English language (Sartori et al. 2016; Dey and Husemann 2018a, b; Harms and Dupérré 2018; Monod et al. 2019; Henningsen et al. 2020; Simoes et al. 2021; Zahiri et al. 2021). We performed a detailed inventory throughout the collection. As a part of this inventory, the entire collection was thoroughly searched and types were transferred to a separate, specially protected type collection. All type specimens were compared with the original descriptions, photographed, databased and relabeled, if required. The original high-quality photographs are available online through the FUNDus! Portal of the University of Hamburg (https://www.fundus.uni-hamburg.de/). The complete data of the type material is provided in two table formats: taxa format (Suppl. material 1: Table S1) and specimen format (Suppl. material 2: Table S2). Our first Lepidoptera catalogue (Zahiri et al. 2021) on the superfamilies Hepialoidea, Cossoidea and Zygaenoidea has resulted in an updated list of 259 as “type” labelled specimens belonging to 13 valid species, 34 subspecies (16 valid and 18 invalid names), and 36 names of invalid infrasubspecific rank (based on the ICZN code).

Material and methods

Photography

All images were taken with a custom-made digital imaging system (DUN Inc.) with stacking capabilities. The images where either taken with a Canon EOS 5DSR with a 65 mm or 100 mm lens and stacked with Zerene Stacker (PMax algorithm) or with a Canon EOS 6D with a 55 mm or 65 mm lens and stacked with Helicon Focus 5.3.

Taxonomy

All taxonomic changes were made after studying the relevant original descriptions of the type material. Synonyms that have already been proposed are listed only if discussed and the type material was examined. Classification used herein follows van Nieukerken et al. (2011). The higher taxonomy of the families Papilionidae, Hesperiidae, Pieridae, Lycaenidae, Riodinidae, and Nymphalidae, as well as the classification below the family level presented here, are based on a combination of recent comprehensive morphological and molecular studies (Paulus et al. 1996; Wahlberg et al. 2009; Heikkilä et al. 2011; Kawahara et al. 2014), as well as the updated checklist of the European butterflies (Wiemers et al. 2018). Finally, like almost all major Lepidoptera databases and catalogues, we did not follow the principle of gender agreement in species names (Sommerer 2002). The recommendation not to follow the articles 31.2 and 34.2 of the ICZN code was adopted at the General Meeting of the Societas Europaea Lepidopterologica (SEL) in a resolution proposed at the 13th European Congress of Lepidopterology in Korsør (Denmark) on June 4, 2002 (van Nieukerken 2019).

Type material

Type localities are cited in their original spelling or with additional remarks (in square brackets), if required. All data was transcribed verbatim as appearing on the respective labels. For unreadable data a question mark (?) was used. Double slash (//) separates data on different labels, a single slash (/) separates lines within each label, and semi-colon (;) separates different specimens. Additional comments are placed in square brackets.

The ICZN (1999) terminologies (e.g., holotype, syntype, lectotype, neotype, paratype, paralectotype, allotype, cotype etc.) are available at the ICZN website (http://iczn.org/content/Glossary) and ICZN articles cited in the text can be found below:

Infrasubspecific taxon - a taxon at lower rank than that of subspecies. The names of such taxa are not regulated by the Code.

Article 45.6.1. - A name is infrasubspecific if its author expressly gave it infrasubspecific rank, or if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity;

Article 45.6.2. - A name is deemed to be infrasubspecific if its author used one of the terms “aberration”, “ab.” or “morph”.

Article 45.6.3. - A name is deemed to be infrasubspecific if it was first published after 1960 and the author expressly used one of the terms “variety” or “form” (including use of the terms “var.”, “forma”, “v.” and “f.”);

Article 45.6.4. - A name is subspecific if first published before 1961 and its author expressly used one of the terms “variety” or “form” (including use of the terms “var.”, “forma”, “v.” and “f.”), unless its author also expressly gave it infrasubspecific rank, or the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, in which case it is infrasubspecific.

Type specimens deposited at the Zoological Museum Hamburg (ZMH)

Superfamily Papilionoidea Latreille, 1802

Papilionoidea (butterflies), currently containing seven families with approximately 18,800 described species worldwide (van Nieukerken et al. 2011), is one of the well-studied lepidopteran groups. The family Nymphalidae (Brush-footed butterflies) with 6,152 described species is the most diverse taxonomic group followed by Lycaenidae (Blues and coppers) with 5,201 species, Hesperiidae (Skippers) with 4,113 species, Riodinidae (Metalmark butterflies) with 1,532 species, Pieridae (Whites or Yellow-whites) with 1,164 species, Papilionidae (Swallotails) with 570 species and Hedylidae with only 36 species (van Nieukerken et al. 2011).

Family Papilionidae Latreille, 1802

Subfamily Parnassiinae Duponchel, 1835

tridentina Dannehl, 1925

Original combination

Parnassius apollo L. (tridentina Dannehl)” Dannehl, 1925 Ent. Z. 39: 6.

Current combination

Parnassius apollo tridentina Dannehl, 1925.

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 61579–61580) (Fig. 1). “Teriol. Mer. / Sum. alp. / Palon / 1.7.24 Ibor [?] / Dannehl” // “Cotype” // “1.7. 1924 / Mt. Palon / Südtirol / Dannehl” // “f. tridentina Dhl.” // “Mus. Altona comm / Eing Nr 3-65” // “ZMH 61579”; “10.6” // “Dannehl / Teriol. mer. / Alp. vai. / Mt. Palon / 1400 m.” // “Cotype” // “10.6 / Mt. Palon / Südtirol / Dannehl” // “Mus. Altona comm / Eing Nr 3-65” // “ZMH 61580”.

Type locality

Italy: “Lokalrasse aus den östlichen Teilen der Trientiner (Sarcataler) Alpen; westl. Etschufer südlich Trient. 1200–1600 m.”

mendolensis Dannehl, 1925

Original combination

Parnassius apollo L. (mendolensis Dannehl).” Dannehl, 1925 Ent. Z. 39: 6.

Current combination

Parnassius apollo mendolensis Dannehl, 1925.

Current status

Valid subspecies.

Type material

Syntypes 1♂1♀ (ZMH 61581–61582) (Fig. 2). “mendolensis ♀” // “Teriol mer. / Mendel / 24. juni / coll. F. Dannehl” // “Cotype” // “Mus. Altona comm / Eing Nr 3-65” // “ZMH 61581”; “mendolensis ♂” // “Teriol mer. / Mendel / 3.Juni.22 / coll. F. Dannehl” // “Cotype” // “Cotype” // “Mus. Altona comm / Eing Nr 3-65” // “ZMH 61582”.

Type locality

Italy: “Fliegt auf den oberen der Mendelstraße bei Bozen und in den meisten östlichen Taleinschnitten des Mt. Roen, 500–1800 m.”

sevensis Kesenheimer, 1921

Original combination

Parn. apollo Linné, nov. subsp. sevensis Ksh.” Kesenheimer, 1921 Ent. Z. 34: 63.

Current combination

Parnassius apollo sevensis Kesenheimer, 1921.

Current status

Junior subjective synonym of Parnassius apollo meridionalis Pagenstecher, 1909.

Type material

Syntypes 1♂1♀ (ZMH 61583–615584) (Fig. 3). “Cotype” // “apollo / sevensis / Cotype ♀ / 19.7.14 / Aalfeld – Sternsee / H.W. Kesenheimer” // “Cotype” // “Sammlung / G. Warnecke / Eing Nr. 5, 1949” // “ZMH 61583”; “Sammlung / G. Warnecke / Eing Nr 51949” // “apollo sevensis / Cotype ♂ / 19.3.14 / Aalfeld - Sternsee / H.W. Kesenheimer” // “Cotype” // “Cotype” // “ZMH 61584”.

Type locality

France: “von Sewen im Masmünstertal in den Vogesen”.

kricheldorffi Eisner, 1928

Original combination

Parnassius apollo, forma kricheldorffi (m.)” Eisner, 1928 Int. Ent. Z. 22: 309.

Current combination

Parnassius apollo kricheldorffi Eisner, 1928.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 61585) (Fig. 4). “Asturien / Picos de Europa / 1800–2000 m. / 20. –28.VII.1928 / leg. A. Kricheldorff” // “v. Kricheldorfi Eisn.” // “Co-Type / e Collection / Bang Haas” // “Cotype” // “Parnassius v. / Kricheldorffi Eis. / det. Eisner” // “Sammlung / G Warnecke / Eing Nr.5. 1949” // “ZMH 61585”.

Type locality

Spain: “Asturien, Picos de Europa, 1800–2000 m”.

Remarks

Eisner (1928) proposed this name as a form of P. apollo (Linnaeus, 1758). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and the author expressly used the terms “variety” or “form”.

ab. praetexta Gelpke, 1929

Original combination

Parnassius apollo L. f. valesiacus ab. praetexta m.” Gelpke, 1929 Int. Ent. Z. 23: 123.

Current combination

Parnassius apollo valesiacus ab. praetexta Gelpke, 1929.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♀ (ZMH 61586) (Fig. 5). “Type / P. apollo L: valesiacus / Fruhst. / ab. praetexta Gelpke / I. E. Z. Guben 23 Jahrg 1929 / S. 122 [illegible]” // “13.7. 1927 / Lötschental / ob Wiler / Wallis 1550 m” // “Type” // “ab. praetexta Gelpke” // “Mus. Altona comm / Eing. Nr. 3-65 // ZMH 61586”.

Original locality

Switzerland: Swiss Alps. “oberhalb Wiler”.

Remarks

Gelpke (1929) proposed this name as an aberration of P. a. valesiacus Fruhstorfer, 1906. Therefore, as stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (if the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable.

ab. uniformis Stauder, 1922

Original combination

Parnassius apollo ab. uniformis” Stauder, 1922 Soc. Ent. 36: 10 (Note)

Current combination

Parnassius apollo ab. uniformis Stauder, 1922.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 61587) (Fig. 6). “Unter-Italien / Stauder 1920” // “uniformis Stdr / Type” // “ab. uniformis Stauder / Type” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61587”.

Original locality

Italy: “Unteritalien”.

Remarks

Stauder (1922) proposed this name as an aberration of P. apollo (Linnaeus, 1758). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph”) and is hence unavailable.

f. bispupillata Turati, 1918

Original combination

Parnassius apollo pumilus Stich. forma bispupillata” Turati, 1918 Atti. Soc. Ital. Sci. Nat. 57: 40.

Current combination

Parnassius apollo pumilus f. bispupillata Turati, 1918.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Cotype” 1♂ (ZMH 61588) (Fig. 7). “Calabrien / [illegible], 1800 m / Stauder, 5.7.20” // “bispupillata” // “bispupillata” // “Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61588”.

Original locality

Italy: Calabria.

Remarks

Turati (1918) proposed this name as a form of P. a. pumilus Stichel, 1906. Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (if the author expressly gave it infrasubspecific rank) and is hence unavailable.

f. appendiculata Turati, 1918

Original combination

Parnassius apollo pumilus Stich. forma appendiculata (f. nuova)” Turati, 1918 Atti. Soc. Ital. Sci. Nat. 57: 41.

Current combination

Parnassius apollo pumilus f. appendiculata Turati, 1918.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Cotype” 1♂1♀ (ZMH 61589–61590) (Fig. 8). “Calabrien / Aspermonte, 1800 m / Stauder 5.7.1920” // “flavomaculata / appendiculata” // “flavomaculata / appendiculata” // “Sammlung / G. Warnecke / Eing. Nr.5, 1949” // “Cotype” // “ZMH 61589”; “Calabrien / Aspromonte, 1800 m / Stauder 5.7.1920” // “nigricans / appendiculata” // “♀ nigricans / appendiculata” // “Sammlung / G. Warnecke / Eing. Nr.5, 1949” // “Cotype” // “ZMH 61590”.

Original locality

Italy: Calabria.

Remarks

Turati (1918) proposed this name as a form of P. a. pumilus Stichel, 1906. Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable.

ab. lacrimans Marschner, 1911

Original combination

Parnassius apollo L. ab. lacrimans” Marschner, 1911 Dt. Ent. Z. Iris 25: 131.

Current combination

Parnassius apollo ab. lacrimans Marschner, 1911.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 61591) (Fig. 9). “Schweizer Jura / Bözinger Berg / Juli 1919 / e coll Marschner / coll. Dr. Gelpke / 1942” // “Type / Parn. apollo L. / f. lacrimans / Marschner / Collect. H. Marschner” // “ab. lacrimans Marschn.” // “Typen” // “Mus Altona comm. / Eing Nr. 3 05” // “ZMH 61591”.

Original locality

Switzerland: “schweizer Jura”.

Remarks

Marschner (1911) proposed this name as an aberration of P. apollo (Linnaeus, 1758). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph”) and is hence unavailable.

oenipontanus Stauder & Felkel, 1921

Original combination

Parnassius apollo oenipontanus, n. subsp.” Stauder & Felkel, 1921 Ent. Anz. 1: 123–124.

Current combination

Parnassius apollo oenipontanus Stauder & Felkel, 1921.

Current status

Valid subspecies.

Type material

Syntypes 10♂♂ (ZMH 61592–61601) (Fig. 10). “e.l. / 9./6.1924 / [illegible] / Innsbruck” // “Coll. Janiter / Leihgabe der / Jungiusgesellschaft / Eing Nr. 13, 1958” // “ZMH 61592”; “Type” // “Innsbruck / 10.6.1913 / J. Felkel” // “oenipontanus Stder / Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61593”; “Type” // “Innsbruck Stilltal” // “Silltal / 7.6.1921 / J. Felkel” // “oenipontanus Stder / Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61594”; “Cotype” // “oenipontanus / Stauder – Felkel / Innsbruk” // “Silltal / 23.5.1921 / J. Felkel” // “Coll. G. Warnecke / Eing. Nr. 11,1983” // “ZMH 61595”; “Type” // “Silltal / 19.5.1921 / J. Felkel” / “oenipontanus / Stauder – Felkel” / “Innsbruck / Silltal” // “oenipontanus Stder / Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61596”; “Cotype” // “Innsbruck / gebirge / 21/6/1914 / J. Felkel leg.” // “oenipontanus Stder / Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61597”; “Type” // Innsbruck / Silltal” // “Silltal / 19.5.1921 / J. Felkel” // “oenipontanus Stauder / Cotype” // “Summlng / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61598”; “Type” // “H. Stauder / Nordtirol / Patsch / 11/6/1920” // “oenipontanus Stauder / Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61599”; “Type” // Innsbruck / 2/6/1914 / J. Felkel leg.” // “subsp. oenipontanus / Stauder / Cotype” // “Summlng / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61600”; “Type” // “Innsbruck / 23.6.1913 / J. Felkel” // “oenipontanus Stder / Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61601”.

Type locality

Austria: “Silltal von Wilten bei Innsbruck bis Patsch, Tirol” [Sill Valley of Wilten near Innsbruck to Patsch, Tirol].

serpentinicus Mayer, 1925

Original combination

Parn. apollo v. serpentinicus, (Rbl. i. 1.) Mayer, subspecies nova” Z. des Öster. Ent.-Ver. 10: 2–4.

Current combination

Parnassius apollo serpentinicus Mayer, 1925.

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 824705–824706) (Fig. 11). “Cotype” // “Burgland / Rötlschlg. / Juli 24 / Mayer” // “Sammlung / G. Warnecke / Eing. Nr 5 , 1949” // “ZMH 824705”; “Cotype” // “Burgland / Stuben / Juli 24 / L. Mayer” // “Serpentinicus Mayer” // “Sammlung / G. Warnecke / Eing. Nr 5 , 1949” // “ZMH 824706”.

Type locality

Austria: “Auf einem Serpentingebirge an der ungarischen Grenze” [on a Serpentin mountain at the Hungarian Border].

ab. marschneri Bryk, 1911

Original combination

Parnassius apollo ab. marschneri” Bryk, 1911 Int. Ent. Z. 5: 161.

Current combination

Parnassius apollo ab. marschneri Bryk, 1911.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 824719) (Fig. 12). “Nordeuropa / Stockholm / Umb. / coll. Dr. Gelpke” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “Type” // “Abgebildet / Bryk” // “Internat. / Entomolog. / Zeitschrift / Guben / V. Jahrg. No. 20” // “ZMH 824719”.

Original locality

Not indicated. Probably Stockholm [Sweden].

Remarks

Bryk (1911) proposed this name as an aberration of P. apollo Linnaeus, 1758, therefore, as stated by article 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

wenzeli Bryk, 1921

Original combination

Parnassius apollo v. Wenzeli” Bryk, 1921 Ent. Tidskr. 42: 117.

Current combination

Parnassius apollo wenzeli Bryk, 1921.

Current status

Valid subspecies.

Type material

Synype 1♂ (ZMH 824721) (Fig. 13). “Cotype” // “Silltal / 19.5.1921 / J. Felkel” // “Coll. G. Warnecke / Eing. Nr. 11 1963” // “ZMH 824721”.

Type locality

“Innsbruck (Patsch)”.

Remarks

Bryk (1921) proposed this name as a variety of P. apollo Linnaeus, 1758. According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”.

gabrieli Bryk, 1934

Original combination

P. nomion ssp. gabrieli (subsp. nova)” Bryk, 1934 Parnassiana 3: 28.

Current combination

Parnassius nomion gabrieli Bryk, 1934.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 61602) (Fig. 14). “Co-Type / Collection / Kotzsch” // “Kukunor / Geb. Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Parnassius / nomion / gabrieli ♂ / Bryk” // “Coll. G. Warnecke / Eing. Nr. 11 1963” // ZMH 61602”.

Type locality

China: “Burchan-Buddha-Kette” [Burchan-Buddha-Chain].

chingamensis Bryk & Eisner, 1932

Original combination

P. nomion F.d.W. chingamensis subsp. nova” Bryk & Eisner, 1932 Parnassiana 2: 26.

Current combination

Parnassius nomion chingamensis Bryk & Eisner, 1932.

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 61604–61605) (Fig. 15). “Parnassius / chinganen- / sis ♂ / C. B.-L / Chingan – Feb. / Ost- Mongolei / lec 7.” // “Coll. G. Warnecke / Eing. Nr. 11. 1963” // “ZMH 61604”; “Mandschur.sept.occ. / Buchalu / Chingan mont. sept. / 1100 m. Juli” // “chinganensis” // “Staudinger / Bang-Haas / Blasewitz” // “Public. Bryk & Eisner / Parnass. II, 2, 1932, 26 / als 1” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61605”.

Type locality

China: “Buchatu, Gudschal-Tal, Chingan mont. sept. 100 m.” [Buchatu, Gudschal-Valley, Chingan mont. sept. 100 m].

Remarks

This appears to be an erroneous original spelling, which was corrected by Eisner (1935) to “chinganensis”. However, Ackery (1973) treats this emendation as an unjustified emendation according to ICZN article 33.2.3 (ICZN 1999) and not as a justified emendation.

tsinlingensis Bryk & Eisner, 1932

Original combination

P. nomion F.d.W. tsinglingensis subsp. nova” Bryk & Eisner, 1932 Parnassiana 2: 26.

Current combination

Parnassius nomion tsinlingensis Bryk & Eisner, 1932.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 61624) (Fig. 16). “Prov. Schensi / Tapaischan [?] / Tsinling schan / 3000 m 20.6.” // “Co-Type / e Collection / Bang Haas” // “subsp. tsinglingensis” // “Public. Bryk ü. Eisner / Parnass. II, 2, 1932 / p. 26” // Sammlung / G. Warnecke / Eing. Nr.5, 1949” // “ZMH 61624”.

Type locality

China: “Schensi, Tapaischan, Tsinlingshan, 3000 m. VI.; Tsingschui, Liu-pinschan, 2000 m.”

nominulus Staudinger, 1894

Original combination

Parnassius Nomion var. Nominulus” Staudinger, 1894 Dt. Ent. Z. Iris 7(2): 241.

Current combination

Parnassius nomion nominulus Staudinger, 1894.

Current status

Valid subspecies.

Type material

Type? 1♂1♀ (ZMH 824717–824718) (Fig. 17). “Irkutsk / Sibirien / A.M. Schmidt / Inh. M. Göttmann” // “Parn. nomion Hb. / Typus?” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 824717”; “Kentei-Geb.” // “Gebr. Dörries leg. 1893 / Erworb. e. coll. / Dörries sen. 1917” // “Nomion typ., [illegible]” // “Sig-G Warnecke / Eing. Nr 5 1949” // “Katalog Nr. / L. 604” // “ZMH 824718”.

Type locality

Russia: “Ost- Sajan -Gebiet (südwestlich von Irkutsk, an der Grenze der chinesischen Mongolei gelegen)” [East-Sajan region (southwest of Irkutsk, at the border of Chinese Mongolia)].

Remarks

Staudinger (1894) proposed this name as a variety of P. nomion Fischer & Waldheim, 1823. According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”. We believe that these two specimens belong to ssp. nominulus Staudinger, 1894. The year on one of the labels (1893) supports this. The question mark on the label (“Typus?”) shows uncertainty about whether it is a type. In the original description, Staudinger says: “... bemerke ich, dass die kleineren Stücke aus dem Kentei-Gebirge den grösseren var. Nominulus fast gleichkommen.“ […I realize that the smaller specimens from the Kentei-Mountains are very similar to the larger var. nominulus].

frivaldszkyi Bang-Haas, 1928

Original combination

Parnassius szechenyii frivaldszkyi O. B.- Haas, subsp. nov.” Bang-Haas, 1928 Ent. Z. 42: 60.

Current combination

Parnassius szechenyii frivaldszkyi Bang-Haas, 1928.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 61603) (Fig. 18). “Kansu occ. / Liang-tschou / Richthofen mont. mar. / 2500 m Juli” // “Co-Type / Haas” // “129” // “szechenyii / frivaldszkyi” // “Mus Altona / Eing Nr 9-64” // “ZMH 61603”.

Type locality

“China sept. occ.: Kansu, Richthofen mont., südwestl. Uang-tschou”.

ab. privatus Kotzsch, 1936

Original combination

Parnassius autocrator Avinoff ab. privatus” Kotzsch, 1936 Ent. Rdsch. 54: 44.

Current combination

Parnassius autocrator ab. privatus Kotzsch, 1936.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Cotype” 1♂ (ZMH 61606) (Fig. 19). “Hindukusch / Chodja-Mahomed / Geröllzone / 3800– 4000 m 25.VII.-10.VIII. / leg. H. & E. Kotzsch” // “Co-Type / e Collection / Kotzsch” // “Mus Altona / Eing 9-64” // “ZMH 61606”.

Original locality

Afghanistan: “Hindukusch, Chodja-Mahomed, Geröllzone, 3800– 4000 m.”

Remarks

Kotzsch (1936) proposed this name as an aberration of P. autocrator Avinoff, 1913. Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph”) and is hence unavailable.

lictor Bryk & Eisner, 1937

Original combination

P. orleans Oberth. subsp. lictor (subsp. nova)” Bryk & Eisner, 1937 Parnassiana 4: 58.

Current combination

Parnassius orleans lictor Bryk & Eisner, 1937.

Current status

Valid subspecies.

Type material

Synypes 1♂1♀ (ZMH 61607–61608) (Fig. 20). “Kukunor/ Geb. Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Parnassius / orleans / lictor ♂ / Br. u. Eis.” // “Co-Type / e Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61607”; “Kukunor/ Geb. Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Parnassius / orleans / lictor ♀ / Br. u. Eis.” // “Co-Type / e Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61608”.

Type locality

China: Kukunor, “Burchan-Buddha, Nomohun Paß”.

buddha Bryk & Eisner, 1937

Original combination

P. mercurius subsp. buddha (subsp. nova)” Bryk & Eisner, 1937 Parnassiana 4: 58.

Current combination

Parnassius jacquemontii buddha Bryk & Eisner, 1937.

Current status

Valid subspecies.

Type material

Syntypes 1♂1♀ (ZMH 61619–61620) (Fig. 21). “Co-Type / e Collection / Kotzsch” // “Kukunor/ Geb. Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Parnassius / mercurius / buddha ♀ / Br. U. Eis.” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61619”; “Kukunor/ Geb. Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Co-Type / e Collection / Kotzsch” // “Parnassius / mercurius / buddha ♂ / Br. U. Eis.” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61620”.

Type locality

China: “Burchan-Buddha-Kette, Nomohun Paß, 4000 m.”

Remarks

Parnassius mercurius Grum-Grshimailo, 1891 is currently considered a subspecies of P. jacquemontii Boisduval, 1836.

tripicta Sheljuzhko, 1928

Original combination

Parnassius bremeri amgunensis” 7. ab. tripicta (nov.)” Sheljuzhko, 1928 Mitt. Münch. Ent. Ges. 18: 7.

Current combination

Parnassius bremeri amgunensis ab. tripicta Sheljuzhko, 1928.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Cotype” 1♂ (ZMH 61621) (Fig. 22). “Mandschur sept. occ. / Buchalu / Chingan mont. Sept. / 1100 m. Juli” // “Cotype” // “Co-Type / e collection / Bang-Haas” // “Bremeri solonensis / ab. tripicta u. flavicans” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61621”.

Original locality

Not indicated. Probably “China, Chingan mont sept., Buchalu.”

Remarks

Sheljuzhko (1928) proposed this name as an aberration of P. b. amgunensis Sheljuzhko, 1928. Therefore, as stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable.

melaniticus Bang-Haas, 1915

Original combination

Parnassius actius melaniticus O. B.-H., nov. var.,” Bang-Haas, 1915 Dt. Ent. Z. Iris 29: 174, t. 5, f. 21.

Current combination

Parnassius actius melaniticus Bang-Haas, 1915.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 833442) (Fig. 23). “Co.Type” // “♂ P. actius / melaniticus” // “Kutscha mont. / Thianschan or / Juli 3500 m.” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 833442”.

Type locality

China: “Tianschan or. mer., Kourgak Taon, Kuldscha Gebirge, nordwestlicher Karaschahr”.

Remarks

Bang-Haas (1915) proposed this name as a variety of P. actius (Eversmann, 1843). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”.

wernickei Kotzsch, 1936

Original combination

Parnassius charltonius Gray subsp. wernickei subsp. nova.” Kotzsch, 1936 Parnassiana 4: 6.

Current combination

Parnassius charltonius wernickei Kotzsch, 1936.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 61622) (Fig. 24). “Nord-Ost-Hindukusch / Nuksan-Pass Nordseite / Alpenwiesenzone / 3500–3800 m Mitte Juli / leg. H. & E. Kotzsch” // “Co-Type / e Collection / Kotzsch” // “983” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 61622”.

Type locality

Afghanistan: “Ost-Hindukusch: Nuksan-Paß, Alpenwiesenzone 3500 bis 4000 m.”.

inopinatus Kotzsch, 1940

Original combination

Parnassius inopinatus” Kotzsch, 1940 Ent. Z. 54: 17.

Current combination

Parnassius inopinatus Kotzsch, 1940.

Current status

Valid species.

Type material

Syntypes 6♂♂ (ZMH 61609–61612, ZMH 824997–824998) (Fig. 25). “Nordwest-Afganistan / Firuskuhi-Mont / Alpenwiesenzone / 2800–3000 m 20.IV.-15.VII. / Exp. Wernicke” // “Parnassius / inopinatus / f Kotzsch” // “Co-Type / Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61609”; “Nordwest-Afganistan / Firuskuhi-Mont / Alpenwiesenzone / 2800–3000 m 20.IV.-15.VII. / Exp. Wernicke” // “Co-Type / Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61610”; “Nordwest-Afganistan / Firuskuhi-Mont / Alpenwiesenzone / 2800–3000 m 20.IV.-15.VII. / Exp. Wernicke” // “Co-Type / Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11 1963” // “ZMH 61611“; “Nordwest-Afganistan / Firuskuhi-Mont / Alpenwiesenzone / 2800–3000 m 20.IV.-15.VII. / Exp. Wernicke” // “Parnassius / inopinatus / m Kotzsch” // “Co-Type / Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61612”; “Nordwest-Afghanistan / Firuskuhi-Mont / Alpenwiesenzone / 2800–3000 m 20.IV.-15.VII. / Exp. Wernicke” // “Co-Type / e Collection / Kotzsch” // “981” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824997”; “Nordwest-Afghanistan / Firuskuhi-Mont / Alpenwiesenzone / 2800–3000 m 20.IV.-15.VII. / Exp. Wernicke” // “Co-Type / e Collection / Kotzsch” // “980” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824998”.

Type locality

“Nordwest-Afghanistan, Firuskuhi-Kette, 2800–3000 m.”.

gigas Kotzsch, 1932

Original combination

Parnassius imperator gigas ssp. n.” Kotzsch, 1932 Ent. Z. 45: 267.

Current combination

Parnassius imperator gigas Kotzsch, 1932.

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 61613–61614) (Fig. 26). “Sincheng / Ost-Sikingshan Gebg. / Juli 2800 m.” // “Parnassius / imperator / gigas f / Her.” // “Cotype” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61613”; “Sincheng / Ost-Sikingshan Gebg. / Juli 2800 m.” // “Parnassius / imperator / gigas m / Her.” // “Cotype” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61614”.

Type locality

China: “vom Ostabhange des Sikingshan Gebirges – Sincheng etwa 2800 m.” [Eastern slopes of the Sikingshan Mountains – Sincheng, ca. 2800 m].

clorinda Kolar & Querci, 1937

Original combination

Parnassius mnemosyne L. v. clorinda” Kolar & Querci, 1937 Lambillionea 5: 99.

Current combination

Parnassius mnemosyne clorinda Kolar & Querci, 1937.

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 824701–824702) (Fig. 27). “Type” // “Macedonia / Olympus, 4500’ / June 16, 1935 / Romei” // “Balkan / coll. Dr. Gelpke” // “Mus Altona / Eing. Nr. 9 – 64” // “ZMH 824701”; “Typen” // “Macedonia / Olympus, 4500’ / June 19, 1935 / Romei” // “Balkan / coll. Dr. Gelpke” // “v. clorinda Querci” // “P. Mnemosyne / - Clorinda Querci / Type abgebildet / in Lambillionea” // “Type” // “Mus Altona / Eing Nr 9-64” // “ZMH 824702”.

Type locality

Macedonia: Olympus.

Remarks

Kolar and Querci (1937) proposed this name as a variety of P. mnemosyne L. According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”.

rjabovi Sheljuzhko, 1935

Original combination

Parnassius mnemosyne rjabovi” Sheljuzhko, 1935 Zs. Öst. Ent. Ver. 20: 22.

Current combination

Parnassius mnemosyne rjabovi Sheljuzhko, 1935.

Current status

Junior subjective synonym of Parnassius mnemosyne caucasia Verity, 1911.

Original material

Labelled as “Autotypes” 1♂1♀ (ZMH 824703–824704) (Fig. 28). “autotypus ♀ / rjabovi Shel.” // “Kleinasien / Armenia / Daratshitshag / 7.VII.1939 / B. Tkatshukov leg. / coll. Dr. Gelpke” // “Mus Altona / Eing Nr 9-64” // “ZMH 824703”; “autotypus ♂ / rjabovi Shel.” // “Kleinasien / Armenia / Daratshitshag / 7.VII.1939 / B. Tkatshukov leg. / coll. Dr. Gelpke” // “v. rjabovi Shelj.” // “Mus Altona / Eing Nr 9-64” // “ZMH 824704”.

Original locality

Armenia: Daratshitshag.

Remarks

The year on the specimen labels (1939) indicate specimens were collected four years after description (1935) from the same location (i.e., Armenia: Daratshitshag) the original holotype material was collected. As a result, the label of the specimens treated as “autotypus” by Warnecke is erroneous. Autotypes do not exist in taxonomy. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these two specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.

comitis Bryk, 1912

Original combination

Parnassius mnemosyne var. comitis” Bryk, 1912 Soc. Ent. 27: 39.

Current combination

Parnassius mnemosyne comitis Bryk, 1912.

Current status

Junior subjective synonym of nominotypical Parnassius mnemosyne (Linnaeus, 1758).

Original material

Labelled as “Cotypen” 1♂ (ZMH 824707) (Fig. 29). “817” // “Italia central. / Majella / 21.VII.1925 / coll. F. Dannehl” // “Italien / coll. Dr. Gelpke” // “Cotypen” // “v. majellensis Dhl.” // “v. comitis Bryk.” // “Mus. Altona comm. / Eing. Nr. 3 65” // “ZMH 824707”.

Original locality

Italy: “aus der Majella Gruppe”.

Remarks

Bryk (1912) proposed this name as a variety of P. mnemosyne (Linnaeus, 1758). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”. The year on the specimen label (1925) indicates specimen was collected 13 years after description (1912) from the same location (i.e., Majella Mountains, Italy) the original holotype material was collected. As a result, the label of the specimen treated as “Cotypen” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, this specimen is erroneously labelled as type, should be annotated as “Not a Type” and is treated here as “Non-type” specimen.

gigantea Staudinger, 1886

Original combination

Parn[assius] mnemosyne L. var. gigantea” Staudinger, 1886 Stett. Ent. Ztg. 47 (4–6): 197.

Current combination

Parnassius mnemosyne gigantea Staudinger, 1886.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 824713) (Fig. 30). “v. gigantea / 1/86 ♂ / Tary. / Ob. Turk. / “Warnecke” // “Tary Ob. Türkestan” // “♂ Parn. Mnemosyne / v. gigantea / E.W. BETH / Hamburg. den Haag” // “Coll. H. Rödinger / Eing. Nr. 30, 1952 / Katalog Nr. L / 6133” // “ZMH 824713”.

Type locality

Russian Turkestan.

Remarks

Staudinger (1886) proposed this name as a variety of P. mnemosyne (Linnaeus, 1758). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”.

beicki Bryk & Eisner, 1932

Original combination

P[arnassius] epaphus beicki subsp. nova (O. Bang-Haas i. l.)” Bryk & Eisner, 1932 Parnassiana 2: 25, 42.

Current combination

Parnassius epaphus beicki Bryk & Eisner, 1932.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 824714) (Fig. 31). “Kansu sept occ / Hsining / Nanchan mont. / Tatung / Nadelholzzone / Jüli 2000 m.” // “Co-Type / e Collection Bang-Haas” // “Ep. subsp. Beiki” // “public. v. Bryk ü. Eisner / Parnass. II, 2, 1932 p. 25” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 824714”.

Type locality

China: “Süd Tatung‘sche Berge, Kansu, 3000 m“ [Southern Tatung Mountains, Kansu, 3000 m].

Remarks

Also misspelled beiki in literature.

ab. rubropicta Bang-Haas, 1915

Original combination

Parnassius epaphus ab. rubropicta” Bang-Haas, 1915 Dt. Ent. Z. Iris 29: 95.

Current combination

Parnassius epaphus ab. rubropicta Bang-Haas, 1915.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Co-Type” 1♂ (ZMH 824715) (Fig. 32). “Karakorum mont. / Schahidulla / Chines Tatarei / 4000 m. Ende Juni” // “Co-Type / e Collection / Bang-Haas” // “subtilis / rubropicata” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824715”.

Original locality

Not indicated. Probably China.

Remarks

Bang-Haas (1915) proposed this name as an aberration of P. epaphus Oberthür, 1879. Therefore, as stated by article 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

candidatus Bang-Haas, 1915

Original combination

Parnassius delphius var. candidatus” Bang-Haas, 1915 Dt. Ent. Z. Iris 29 (2/3): 158.

Current combination

Parnassius delphius candidatus Bang-Haas, 1915.

Current status

Junior subjective synonym of the nominotypical Parnassius delphius Eversmann, 1843.

Type material

Syntype 1♂ (ZMH 824716) (Fig. 33). “Chin. Turkestan / Thianschan mont. or. / Juldus Tal / 2500 m. Juli” // “Co-Type / O. Bang-Haas” // “delphiuscaudi- / datus Cotype” // “Slg. Mus. Altona / Eing. nr. 6/1978” // “ZMH 824716”.

Type locality

“Tianschan or.; Juldus (Kuldsca, Ili, Issykkul)”.

Remarks

Bang-Haas (1915) proposed this name as a variety of P. delphius Eversmann, 1843. According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”.

eva Kotzsch, 1936

Original combination

Parnassius delphius Eversm. subsp. eva subsp. nova” Kotzsch, 1936 Parnassiana 4: 5.

Current combination

Parnassius jacobsoni eva Kotzsch, 1936.

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 824723–824724) (Fig. 34). “Hindukusch / Chodja-Mahomed / Geröllzone / 3800.4000 m 25.VII.-10.VIII. / leg. H. & E. Kotzsch” // “Co-Type / e Collection Kotzsch” // “977” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824723”; “Hindukusch / Chodja-Mahomed / Geröllzone / 3800.4000 m 25.VII.-10.VIII. / leg. H. & E. Kotzsch” // “Co-Type / e Collection Kotzsch” // “978” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824724”.

Type locality

“Hindukusch, Chodja-Mahomed” [Afghanistan, Hindukusch, Khoja Mohammad Mts].

Unpublished, unavailable names

minor Frank

Original combination

Parnassius mnemosyne hartmanni ab. minor Frank.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 824708) (Fig. 35). “Nördl. Kalkalpen / Inntaler Berge / Oberaudorf / 1.VI.24 500-700 m / coll. Dr. Gelpke” // “Mus. Altona comm. / Eing. Nr. 3-65” // “Type” // “ab. minor Frank” // “Mus. Altona comm. / Eing Nr. 3 65” // “ZMH 824708”.

Original locality

Nördl. Kalkalpen [Austria].

Remarks

This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable. The current name of the subspecies is Parnassius mnemosyne hartmanni Standfuss, 1888.

nigra Frank

Original combination

Parnassius mnemosyne hartmanni ab. nigra Frank.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 824709) (Fig. 36). “ab. nigra Frank” // “Nördl. Kalkalpen / Chiemgau / Lofer/ 28.V.1920 / coll. Dr. Gelpke” // “Type” // “ab. nigra Frank” // “Mus. Altona comm. / Eing Nr 3 - 65” // “ZMH 824709”.

Original locality

Nördl. Kalkalpen [Austria].

Remarks

This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable. The current name of the subspecies is Parnassius mnemosyne hartmanni Standfuss, 1888.

ab. punctata Frank

Original combination

Parnassius mnemosyne hartmanni ab. punctata Frank.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂, “Cotype” 1♂ (ZMH 824710–824711) (Fig. 37). “Cotypen” // “Nördl. Kalkalpen / Inntaler Berge / Oberaudorf / 1.VI.24 500-700 m / coll. Dr. Gelpke” // “Mus. Altona comm. / Eing. Nr. 3-65” // “ZMH 824710”; “Typen” // “Mus. Altona comm. / Eing. Nr. 3-65” // “Nördl. Kalkalpen / Inntaler Berge / Oberaudorf / 30.V.23 500-700 m / coll. Dr. Gelpke” // “ab. punctate Frank” // “ZMH 824711”.

Original locality

Nördl. Kalkalpen [Austria].

Remarks

This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable. The current name of the subspecies is Parnassius mnemosyne hartmanni Standfuss, 1888.

nigerrima Frank

Original combination

Parnassius mnemosyne hartmanni ab. nigerrima Frank.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 824712) (Fig. 38). “Nördl. Kalkalpen / Inntaler Berge / Oberaudorf / 1.VI.24 500-700 m / coll. Dr. Gelpke” // “Type” // “Mus. Altona comm. / Eing. Nr. 3-65” // “ab. nigerrima Frank” // “ZMH 824712”.

Original locality

Nördl. Kalkalpen [Austria].

Remarks

This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable. The current name of the subspecies is Parnassius mnemosyne hartmanni Standfuss, 1888.

ab. subtusalbipupillata

Original combination

Parnassius apollo ab. subtusalbipupillata.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 824720) (Fig. 39). “Balkan / Bulgaria / Sofia / Mt. Vitosch / 2000 m Juli” // “ab. quincunx Bryk / ab. reniformis O. B.-H. / ab. laticincta Trti.” // “Mus. Altona comm. / Eing. Nr. 3-65” // “Type” // ZMH 824720”.

Original locality

Balkan, Bulgaria.

Remarks

This taxon cannot be found in the literature. As stated by article 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable. The current name of the species is Parnassius apollo (Linnaeus, 1758).

analifasciata Gelpke

Original combination

Parnassius apollo ab. analifasciata Gelpke.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 824722) (Fig. 40). “Süddeutschland / Fränk. Jura / Eichstätt / 12.VII.1921 / coll. Dr. Gelpke” // “Type” // “ab. analifasciata Gelpke” // “ab. splendida Osth. – fasciata st. / ab. magnifica Bryk / ab. subt-bip. Fr. ab. subt-graphica / ab. seitzi Brk. – embryki Bryk.” // “Mus. Altona comm. / Eing. Nr. 3-65” // “ZMH 824722”.

Original locality

South Germany.

Remarks

This taxon cannot be found in the literature. As stated by article 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable. The current name of the species is Parnassius apollo (Linnaeus, 1758).

Subfamily Papilioninae Latreille, 1802

Tribe Papilionini Latreille, 1802

f. unimaculata Warnecke, 1924

Original combination

Papilio xuthus Linnaeus f. unimaculata” Warnecke, 1924 Dt. Ent. Z. Iris 38: 149.

Current combination

Papilio xuthus f. unimaculata Warnecke, 1924.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 61623) (Fig. 41). “Japan” // “Type” / / Sammlung / Fritz Diehl / Eing. Nr. 35, 1950” // “P. xuthus L. / n.f. // unimaculata / Warn.” // “ZMH 61623”.

Original locality

Original publication without location records; label states Japan; according to hand written type catalogue of Warnecke: Ussuri ex Coll. Dörries (67), holotype destroyed in fires in Zool. Mus. Hamburg 1943, 1 specimen with location label Japan labelled as type by Warnecke in Coll. Warnecke, now in ZMH.

Remarks

Warnecke (1924) proposed this name as a form of P. xuthus Linnaeus, 1776. According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. The text of Warnecke (1942) paper unambiguously reveals that he describes an individual form and not a (geographic) subspecies.

Tribe Troidini Talbot, 1939

peruanus Fuchs, 1954

Original combination

Papilio archidamas var. peruanus” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 83–84.

Current combination

Battus polydamas peruanus (Fuchs, 1954).

Current status

Valid subspecies.

Type material

Lectotype 1♂ (ZMH 61615) (Fig. 42) Paralectotypes 2♂1♀ (ZMH 61616–61617, ZMH 824999). “Rio Pampas / Cuesta de Bonbon / 8/1928 / ca. 2000 m / Type” // “HOLOTYPE / Papilio archidamas / var. peruanus / von Fuchs” // “archidamas / S. Rio Pampas / 2000 m” // “Z.I.M. / Hamburg” // “LECTOTYPE ♂ / Papilio archidamas / var. peruanus Fuchs / by G. Lamas, 1978” // “Coll. H. Rödinger / Eing. Nr. 30, 1952 / Katalog Nr. L.” // “Battus archidamas / peruanus von Fuchs / G. Lamas. Det. 1978” // “Battus polydamas / peruanus (Fuchs, 1954) / Lectotype ♂ / Racheli & Pischedda 1987” // “ZMH 61615”; “Rio Pampas / Cuesta de Bonbon / 8/1928 / ca. 2000 m / Type” // “P. archidamas peruanus” // “Z.I.M. Hamburg” // “Battus polydamas / peruanus (Fuchs, 1954) / Paralectotype ♂ / Racheli & Pischedda, 1987” // “ZMH 61616”; “Rio Pampas / Cuesta de Bonbon / 8/1928 / ca. 2000 m / Type” // “Battus polydamas / peruanus (Fuchs, 1954) / Paralectotype ♀ / Racheli & Pischedda, 1987” // “Z.I.M. Hamburg” // “ZMH 61617”; “¿” // “Papilio / archidamas ♂ / var. peruanus / von Fuchs / Rio Pampas / Cuesta de Bonbon / c. 2000 m / 8/28” // “Z.I.M. / Hamburg” // “Type” // “Battus polydamas / peruanus (Fuchs, 1954) / Paralectotype ♂ / Racheli & Pischedda 1987” // “Coll. H. Rödinger / Eing. Nr. 30, 1952 / Katalog Nr. L.” // “ZMH 824999”.

Type locality

[Lectotype] Peru: “Rio Pampas, Cuesta de Bonbon, ca. 2000 m”; lectotype designated by Lamas (1978).

Remarks

Fuchs (1954) proposed this name as a variety of P. archidamas Boisduval, 1836. According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”. Papilio archidamas Boisduval, 1836 is a junior subjective synonym of Battus polydamas (Linnaeus, 1758).

Tribe Leptocircini Kirby, 1896

ab. ornatissima Warnecke, 1924

Original combination

Papilio podalirius L. n. ab. ornatissima Warn.” Warnecke, 1924 Int. Ent. Z. 18: 157.

Current combination

Iphiclides podalirius ab. ornatissima (Warnecke, 1924).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 61618) (Fig. 43). “n. ab. ornatissima / Warnecke. Type” // “Type” // “ältester [?] Name: confluens Ver. / Rhop. Pal. P. 292, t. 57, f. 5, 19ii / vgl Lambillioneaa 1932 p. 223” // “Katalog Nr. / L. 204” // “Slg. G. Warnecke / eing. Nr. 5. 1949” // “ZMH 61618”.

Original locality

Southern Germany.

Remarks

Warnecke, (1924) proposed this name as an aberration of Papilio podalirius Linnaeus, 1758. Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph”) and is hence unavailable.

Unpublished, unavailable names

ab. indistinctus Meinhard

Original combination

Iphiclides feisthamelii f. ornatus ab. indistinctus Meinhard.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 835574) (Fig. 44). “26/5. 1906 / Stuttgart / ex larv.” // “ab. indistinctus / Meinh.” // “ZMH 835574”.

Remarks

This taxon cannot be found in the literature. Probably an aberration of Iphiclides feisthamelii (Duponchel, 1832). Apparently the imago emerged in Stuttgart (Germany), but the larva must have been collected in the Iberian Peninsula (M. Wiemers pers.com.).

Family Pieridae Swainson, 1820

Subfamily Coliadinae Swainson, 1827

crocopepla Warnecke, 1912

Original combination

Colias aurora Esp. ♀ ab. crocopepla” Warnecke, 1912 Soc. Ent. 27: 68.

Current combination

Colias heos ab. crocopepla Warnecke, 1912.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type”1♀ (ZMH 827594) (Fig. 45). “Type” // “Bikienfl.” // “ab. crocopepla / Warn. Soc. Ent. 27 ö. 68” // “L.6” // “sig. G. Warnecke / Eing. Nr. 5, 1949” // “Katalog Nr. / L 1754” // “AMUR-GEBIET: / Gebr. Dörries leg. 1887 / Erworb. e. coll. / Dörries sen. 1917 / (vgl. Mém. Rom. VI, / 1892 p 91)” // “ZMH827594”.

Original locality

Amur [Russia].

Remarks

Papilio aurora Esper, 1783 is a junior primary homonym of Papilio aurora Cramer, [1780]. The former is currently known as Colias heos (Herbst, 1792) and the latter is known as Colotis aurora (Cramer, [1780]). Grieshuber and Lamas (2007) provided a nomenclatural list of all available and unavailable names in the genus Colias and clearly stated that Colias aurora [ab.] crocopepla is infrasubspecific and thus not available name.

ab. rhododactyla Warnecke, 1912

Original combination

Colias aurora Esp. ♀ ab. rhododactyla” Warnecke, 1912 Soc. Ent. 27: 68.

Current combination

Colias heos ab. rhododactyla Warnecke, 1912.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♀ (ZMH 827593) (Fig. 46). “L.39 / Col. aurora / ab. ♀ rhododactyla / Warn.” // “ab. rhododactyla / Warnecke / Soc. Ent. 27, 1912 p. 68” // “Katalog Nr. / L. 1755” // “Tjutjuché / Ost-Siberien” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827593”.

Original locality

Amur [Russia].

Remarks

Papilio aurora Esper, 1783 is a junior primary homonym of Papilio aurora Cramer, [1780]. The former is currently known as Colias heos (Herbst, 1792) and the latter is known as Colotis aurora (Cramer, [1780]). Grieshuber and Lamas (2007) provided a nomenclatural list of all available and unavailable names in the genus Colias and clearly stated that Colias aurora [ab.] rhododactyla is infrasubspecific and thus not available name.

ab. theia Warnecke, 1912

Original combination

Colias aurora Esp. ♀ ab. theia” Warnecke, 1912 Soc. Ent. 27: 68.

Current combination

Colias heos ab. theia Warnecke, 1912.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Lectotype” 1♀ (ZMH 827609) (Fig. 47). “Kentei- / Gebirge” // “Gebr. Dörries leg.1889 / Erworb. E. coll. / Dörries sen. 1917” // “f. theia Warnecke / Die Type ist im Zool. Mus. Hamburg!” // “L. 38 / Lekto-Type / Col. aurora f. theia warn.” // “Katalog Nr. / L 1750” // “Sig. G. Warnecke / Eing. Nr. 5, 1945” // “ZMH 827609”.

Original locality

Amur [Russia].

Remarks

Papilio aurora Esper, 1783 is a junior primary homonym of Papilio aurora Cramer, [1780]. The former is currently known as Colias heos (Herbst, 1792) and the latter is known as Colotis aurora (Cramer, [1780]). Grieshuber and Lamas (2007) provided a nomenclatural list of all available and unavailable names in the genus Colias and clearly stated that Colias aurora [ab.] theia is infrasubspecific and thus not an available name.

f. rebeli Schawerda, 1906

Original combination

Colias myrmidone balcánica f. rebeli” Schawerda, 1906; Verh. Zool. Bot. Ges. Wien 6: 651.

Current combination

Colias caucasica balcanica f. rebeli Schawerda, 1906.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Cotype” 1♀ (ZMH 827619) (Fig. 48). “Orjen / Herzegowina / Dr. Schawerda” // “Sammlung / Fritz Diehl / Eing. Nr. 35,1950” // “rebeli ♀” // “Cotype” // “Cotype” // “L5” // “Katalog Nr. L 1721” // “ZMH 827619”.

Original locality

Bosnia and Herzegovina: “vom Trebević”.

Remarks

Schawerda (1906) proposed this name as a form of C. m. balcanica Rebel, 1901. Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. Colias balcanica Rebel, 1901 is currently regarded as subspecies of C. caucasica Staudinger, 1871. Grieshuber and Lamas (2007) provided a nomenclatural list of all available and unavailable names in the genus Colias and clearly stated that Colias myrmidone balcanica [f.] rebeli is infrasubspecific and thus not available name.

tatarica Bang-Haas, 1915

Original combination

Colias cocandica tatarica O. B.-H. n. var.” Bang-Haas, 1915 Dt. Ent. Z. Iris 29: 98.

Current combination

Colias cocandica tatarica Bang-Haas, 1915.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 833446) (Fig. 49). “Co-Type” // “Chotan mer / Karakorum / Juli” // “VII. / Chotan. mer. / Karakorum / v. Friedrich [?]” // “ZMH 833446”.

Type locality

China: “… am Nordabhange des Karakorum, in den Gebirgen südlich der Stadt Chotan (Chinesische Tatarei)” […around the northern slopes of the Karakorum, in the Mountains south of the town of Chotan (Chinese Tartarei)].

Remarks

Bang-Haas (1915) proposed this name as a variety of C. cocandica Erschoff, 1874. According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”.

aurea Kotzsch, 1936

Original combination

Colias wiskotti aurea subsp. n.” Kotzsch, 1936 Ent. Rdsch. 54(4): 45.

Current combination

Colias wiskotti aurea Kotzsch, 1936.

Current status

Valid subspecies.

Type material

Syntypes 3♂♂ 1♀ (ZMH 827622–827623, ZMH 835580–835581) (Fig. 50). “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H. & E. Kotzsch” // “Wisk. aurea Kotzsch” // “Co-Type / e Collection / Kotzsch” // “L.7” // “Katalog Nr. / L 1793” // “Sig G. Warnecke / Eing. Nr. 5, 1946” // “ZMH 827622”; “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H. & E. Kotzsch” // “Katalog Nr. / L 1794” // “Wiskotti aurea Ktz” // “Co-Type / e Collection / Kotzsch” // “L.8” // “Sig G. Warnecke / Eing. Nr. 5, 1946” // “ZMH 827623”; “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H. & E. Kotzsch” // “Co-Type / e Collection / Kotzsch” // “Wiskotti ♂/ aurea” // “ZMH 835580”; “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H. & E. Kotzsch” // “Colias ♀ / wiskotti / v. aurea” // “Co-Type / e Collection / Kotzsch” // “ZMH 835581”.

Type locality

Afghanistan:“Ost-Hindukusch, Nordseite, Alpenwiesenzone, 3500–3800 m.”

yauta Fuchs, 1928

Original combination

Teriocolias yauta” Fuchs, 1954 Beiträge zur Fauna Perus 4:83.

Current combination

Eurema zelia andina (Fuchs, 1928).

Current status

Junior subjective synonym of Eurema zelia andina (Forbes, 1928).

Type material

Holotype 1♂ (ZMH 825001), Paratypes 8♂♀ (ZMH 825000, ZMH 825002–825008) (Fig. 51). “Teriocolias / yauta ♂ / J. Fuchs / Cordillere von / Ilaula 5.22 / 3000 m” // “Sammlung / W.v. Fuchs / Eing. Nr. 4. 1956” // “Type” // “Paratype / Teriocolias yauta / von Fuchs” // “Teriocolias zelia / andina Forbes / G. Lamas. det. 1978” // “ZMH 825000”; “Llauta / ca. 3000 m / 5/1922 / Type” // “Coll. H. Rödinger / Eing. Nr. 30, 1952” // “Holotype / Teriocolias yauta / von Fuchs” // “Teriocolias zelia / andina Forbes / G. Lamas. det. 1978” // “ZMH 825001”; “Llaüta / ca. 3000 m / 5/1922 / Type” // “Coll. H. Rödinger / Eing. Nr. 30, 1952” // “ZMH 825002”; “Laramate / ca 3400 m / 5/1922 / Type” // “Coll. H. Rödinger / Eing. Nr. 30, 1952” // “ZMH 825003”; “Oberhalb / Aina / ca 3000 m / 8/1924 / Type” // “Coll. H. Rödinger / Eing. Nr. 30, 1952” // “ZMH 825004”; “Llaüta / ca. 3000 m / 5/1922 / Type” // “Coll. H. Rödinger / Eing. Nr. 30, 1952” // “ZMH 825005”; “Type” // Teriocolias / yauta ♀ / v. Fuchs / Cordillere von / Ilaula 5.22 / 3000 m” // “Sammlung / W.v. Fuchs / Eing. Nr. 4. 1956” // “ZMH 825006”; “Type” // “Süd – Peru / Ocana, 2600 m / 7.4.1936.” // “Hbg. – Süd – Peru / Sammelreise 1936. / Eing. Nr. 1, 1937.” // “Teriocolias / yauta ♂ / v. Fuchs” // “Sammlung / W, v Fuchs / Eing. Nr. 4, 1956” // “ZMH 825007”; “Type” // Teriocolias / yauta ♀ / v. Fuchs / Cordillere von / Ilaula 5.22 / 3000 m” // “Sammlung / W.v. Fuchs / Eing. Nr. 4. 1956” // “ZMH 825008”.

Type locality

South Peru. “ … Kordillere von Llauta … auch an anderen Stellen der Kordilleren” ” [Cordillera of Llauta…also other areas in the cordilleras].

Subfamily Dismorphiinae Schatz, 1887

lorkovici Pfeiffer, 1932

Original combination

Leptidea duponcheli Stgr. ssp. n. lorkovici m.” Pfeiffer, 1932 in Osthelder and Pfeiffer 1932, Mitt. Münch. Ent. Ges. 22: 20.

Current combination

Leptidea duponcheli lorkovici Pfeiffer, 1932.

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 827614–827615) (Fig. 52). “Syr. Sept. / Taurus c. / Marasch / 6–900 m V.30 / Einh. Slr. leg.” // “Cotypus / duponcheli / ssp. lorkovici / Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827614”; “Syria s. / Amanus s. / (Düldül Dagh) / Jeschildere / VI 33 / Coll. Pfeiffer / München” // “Cotypus / duponcheli / ssp. lorkovici / Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr 20 - 60” // “ZMH 827615”.

Type locality

Turkey: “Marasch in türkisch Nordsyrien; An den Hängen um M bis 1200 m” [Marasch in the turkish regions of northern Syria. On the slopes around Marasch up to 1200 m].

f. fragilis Dannehl, 1925

Original combination

Leptidea sinapis L. (gen. autumnalis) v. fragilis Dannehl” Dannehl, 1925 Ent. Z. 39: 6.

Current combination

Leptidea sinapis f. fragilis Dannehl, 1925.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Cotype” 1♂ (ZMH 835575) (Fig. 53). “3.10.24” // “Südtirol Terlan / Franz Dannehl” // “fragilis Dhl. ” // “Cotype” // “3.10 1924 / Terlan / Südtirol / Dannehl” // “ZMH 835575”.

Original locality

Italy: “in Südtirol bei Bozen, Fleimstal, Nonstal.”

Remarks

Dannehl (1925) proposed this name as a form of L. sinapis (Linnaeus, 1758). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. The text of Dannehl (1925) paper unambiguously reveals that this form was described as a seasonal form (autumn generation) and not as a subspecies.

ab. simbruina Dannehl, 1927

Original combination

Leptidea sinapis L. ab. simbruina Dhl.” Dannehl, 1927 Mitt. Münch. Ent. Ges. 17: 1.

Current combination

Leptidea sinapis ab. simbruina Dannehl, 1927.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Cotype” 2♂♂ (ZMH 835582–835583) (Fig. 54). “25.6” // “Italia central. / Mti. Simbruini / coll. F. Dannehl” // “simbruina” // “Cotypen” // “25.6. 1927 / Mte Simbruini / Central Italien” // “ZMH 835582”; “7.4. 1927 / Mte Simbruini / Central Italien” // “ZMH 835583”.

Original locality

Italy: “Sabiner und Italy: Simbruiner Berge vom Tal (Campagna) bis auf die Gipfel der angegebenen Gebirge.” [Sabiner and Italy: Simbruin Mountains from the valley (Campagna) up to the top of the stated mountain ranges].

Remarks

Dannehl (1927) proposed this name as an aberration of L. sinapis (Linnaeus, 1758). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph”) and is hence unavailable.

stabiarum Staudinger, 1914

Original combination

Leptidea (Leucophasia) sinapis stabiarum m., nov. subspec.” Staudinger, 1914 Zeit. Wiss. Ins. Biol. 10: 371, t. 11, figs. 5, 6.

Current combination

Leptidea sinapis stabiarum Staudinger, 1914.

Current status

Junior subjective synonym of nominotypical Leptidea sinapis (Linnaeus, 1758).

Type material

Syntypes 3♂♂ (ZMH 827624–827626) (Fig. 55). “Ital. mer. 600 m. / penins. Surrentina / M. Faito 3.6.1920 / H. Stauder legit.” // “Type” // “Sammlung / G Warnecke / Eing Nr 5, 1949” // “ZMH 827624”; “25/6 / Ital. mer. m. / penins. Surrentina / M. Fatio 3.6.1920 / H. Stauder legit.” // “Type” // “Sammlung / G Warnecke / Eing Nr 5, 1949” // “ZMH 827625”; “26/6 / Ital. mer. 700 m. / penins. Surrentina / M. Fatio 3.6.1920 / H. Stauder legit.” // “Type” // “Sinapis stabiarum / Stauder / v. stabiarum.” // “ZMH 827626”.

Type locality

Italy: Monte Faito and S. Angelo [Naples], 700–1400 m.

Subfamily Pierinae Swainson, 1820

italorum Dannehl, 1929

Original combination

Anthocharis euphenoides St. italorum Dhl.” Dannehl, 1929 Mitt. Münch. Ent. Ges. 19: 98.

Current combination

Anthocharis euphenoides italorum Dannehl, 1929.

Current status

Junior subjective synonym of nominotypical Anthocharis euphenoides Staudinger, 1869.

Type material

Syntype 1♂ (ZMH 835584) (Fig. 56). “20.5” // “Apenn. Central. / Mte. Genzana / 1500–2000 m. / coll. F. Dannehl” // “Cotype” // “20.5.1928 / Mte Genzana / Central- Apenninen / 1500–2000 m D.” // “ZMH 835584”.

Type locality

Italy: “… in den Gebieten des Velino, Sirente, Monte Genzana, Monte Paradiso und der Majella. auf Höhen von 1500–2000 m.” [in the mountain ranges of the Velino, Sirente, Monte Genzana, Monte Paradiso and Majella. In altitudes of 1500–2000 m].

paravicinii Stauder, 1915

Original combination

Euchloë belia paravicinii” Stauder, 1915 Dt. Ent. Z. Iris 29: 25.

Current combination

Euchloe ausonia paravicinii Stauder, 1915.

Current status

Junior subjective synonym of Euchloe ausonia melanochloros Röber, 1907.

Type material

Syntype 1♂ (ZMH 827616) (Fig. 57). “Algeria mer. / Djebel Aures / 10.5.1914 / H. Stauder” // “Paravicinii Stauder” // “paravicinii Stauder” // “Type” // “L. 4” // “Katalog Nr. / L. 1180” // “Type” // “Sig. G. Warnecke / Eing. Nr. 5, 1948” // “ZMH 827616”.

Type locality

Algeria: “Djebel Aurès in Südalgerien”.

Remarks

Papilio belia Stoll [1782] is a primary homonym of Papilio belia Linnaeus, 1767 which is currently known as Anthocharis belia.

f. flavide apicata Stauder, 1922

Original combination

Teracolus daira nouna auresiaca g. II. interposita Stdr. f. flavide apicata m. nov.” Stauder, 1922 Mitt. Münch. Ent. Ges. 12: 24–25.

Current combination

Colotis evagore nouna f. flavide apicata (Stauder, 1922).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 7♂♂ (ZMH 827595–827599, ZMH 827601–827602) (Fig. 58). “Type” // “El Kautara / auf VI” // “Auresica gen. II. / interposita Stauder” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827595”; “♂” // “Type” // “Algeria mer. / Djebel Aures / VI. 1913 / H. Stauder” // “f. interposita / 5.VI” // “VI form / ♂ 200” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827597”; “Type” // “El Kautara / auf VI” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827596”; “Type” // “El Kautara / auf VI” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827598”; “♂” // “Type” // “Algeria mer. / Djebel Aures / VI. 1912 / H. Stauder” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827599”; “Type” // “X Djebel Aures / VII. 1912 H. Stauder” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827601”; “♂” // “Type” // “Algeria mer. / Djebel Aures / VI. 1913 / H. Stauder” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827602”.

Original locality

Not indicated. Probably “Algeria mer. / Djebel Aures”.

Remarks

Stauder (1922) proposed this name as a form of T. d. nouna auresiaca interposita Stauder, 1922. Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. Teracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).

f. vernalis alticola auresiaca Stauder, 1913

Original combination

Teracolus daira (= dalia Fldr.) nouna Luc. 1848 (= demagore Fldr.) forma vernalis alticola auresiaca” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290.

Current combination

Colotis evagore nouna f. vernalis alticola auresiaca (Stauder, 1913).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 3♂3♀ (ZMH 827600, ZMH 827603–827607) (Fig. 59). “♂” // “Cotype” // “Algeria mer. / Djebel Aures / V. 1912 / H. Stauder” // “auresiaca” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827600”; “♂” // “Type” // “Algeria mer. / Djebel Aurès / V. 1912 / H. Stauder” // “auresiaca / ♂ Stdr.” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827603”; “♀” // “Type” // “Algeria mer. / Djebel Aurès / V. 1912 / H. Stauder” // “auresiaca Stdr. / ♀” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827604”; “♀” // “Cotype” // “Algeria mer. / Djebel Aurès / V. 1912 / H. Stauder” // “auresiaca / ♀” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827605”; “♀” // “Cotype” // “Algeria mer. / Djebel Aurès / V. 1912 / H. Stauder” // “auresiaca / ♀” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827605”; “♂” // “Type” // “Algeria mer. / Djebel Aurès / V. 1912 / H. Stauder” // “auresiaca Stdr. / ♂” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827607”.

Original locality

Algeria: “in den algerischen Atlas und die nördliche Sahara” [in the Algerian Atlas and the northern Sahara].

Remarks

Stauder (1913) proposed this name as a form of T. d. nouna (Lucas, 1848). Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. Teracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).

f. biformata Stauder, 1913

Original combination

Teracolus daira (= dalia Fldr.) nouna Luc. 1848 (= demagore Fldr.) forma autumnalis biformata m.” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290.

Current combination

Colotis evagore nouna f. biformata (Stauder, 1913).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827608) (Fig. 60). “♂” // “biformata / ♂ Stdr.” // “Type” // “Algeria mer. / Djebel Aurès / V. 1912 / H. Stauder” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827608”.

Original locality

Algeria: “in den algerischen Atlas und die nördliche Sahara” [in the Algerian Atlas and the northern Sahara].

Remarks

Stauder (1913) proposed this name as a form of T. d. nouna (Lucas, 1848). Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. Teracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).

f. aestiva pyroleuca Stauder, 1913

Original combination

Teracolus daira nouna Luc. forma aestiva pyroleuca m.” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290.

Current combination

Colotis evagore nouna f. aestiva pyroleuca (Stauder, 1913).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type and Cotype” 10♂♀ (ZMH 827618, ZMH 827628–827636) (Fig. 61). “♀” // “Djebel Aurès / VII. 1912 Stauder” // “nouna var. / pyroleuca / Stauder” // “TYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827618”; “♀” // “Algeria mer. / Djebel Aurès / VII. 1912 / H. Stauder” // “pyroleuca ♀” // “COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827628”; “♂” // “Algeria mer. / Djebel Aurès / VII. 1912 / H. Stauder” // “Cotype” // “COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827629”; “♂” // “Algeria mer. / Djebel Aurès / VII. 1912 / H. Stauder” // “pyroleuca ♂” // “TYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827630”; “♀” // “Algeria mer. / Djebel Aurès / VII. 1912 / H. Stauder” // “pyroleuca ♀” // “COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827631”; “♂” // “Algeria mer. / Djebel Aurès / VII. 1912 / H. Stauder” // “pyroleuca ♂” // “COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827632”; “♂” // “Algeria mer. / Djebel Aurès / VII. 1912 / H. Stauder” // “pyroleuca ♂ NO” // “COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827633”; “♂” // “COTYPE” // Algeria mer. / El Kaictara [?] / anfg IX 19 / H. Stauder” // “pyroleuca ♂” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827634”; “♀” // “Type” // “Algeria mer. / Djebel Aures / VII. 1912 / H. Stauder” // “pyroleuca ♀” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827635”; “♂” // “TYPE” // “Algeria mer. / Djebel Aurès / VII. 1913 / H. Stauder” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827636”.

Original locality

Algeria: “in den algerischen Atlas und die nördliche Sahara” [in the Algerian Atlas and the northern Sahara].

Remarks

Stauder (1913) proposed this name as a form of T. d. nouna (Lucas, 1848). Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. Teracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).

ab. evagorides Stauder, 1913

Original combination

Teracolus daira nouna Luc. ab. evagorides m.” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290.

Current combination

Colotis evagore nouna ab. evagorides (Stauder, 1913).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Cotype” 2♂♂ (ZMH 827620–827621) (Fig. 62). “♂” // “Algeria mer. / Djebel Aurès / VI. 1913 / H. Stauder” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “COTYPE” // “ZMH 827620”; “♂” // “evagorides / Stdr.” // “Algeria mer. / Djebel Aurès / VII. 1912 / H. Stauder” // “TYPE” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827621”.

Original locality

Algeria: “in den algerischen Atlas und die nördliche Sahara” [in the Algerian Atlas and the northern Sahara].

Remarks

Stauder (1913) proposed this name as an aberation of T. d. nouna (Lucas, 1848). Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. Teracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).

italorum Stauder, 1921

Original combination

Pieris brassicae L. f. n. (g. aest. mer. alticola) italorum” Stauder, 1921 Dt. Ent. Z. Iris 35: 26.

Current combination

Pieris brassicae italorum Stauder, 1921.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 827617) (Fig. 63). “Calabria mer. / Aspromonte 900 m. / 29/6 Polsi [?] 1920 / H. Stauder Legit.” // “Type” // “brassicae var. / italorum Staud.” // “ZMH 827617”.

Type locality

Italy: “i.[talien] c.[entralis] m.[eridionalis], Aspromonte, Montalto, 1800 m.”

elbursina Bytinski-Salz & Brandt, 1937

Original combination

Pieris ergane, Hbn. ssp. elbursina, ssp. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 49: 1–2.

Current combination

Pieris ergane elbursina Bytinski-Salz & Brandt, 1937.

Current status

Valid subspecies.

Type material

Syntypes 3♂♂ 1♀ (ZMH 827610–827613) (Fig. 64). “IRAN / Keredj (1500 m.) / 18/5 1936 / Brandt” // “Cotypus / ergane ssp. / pfeifferi / By.S+Br.” // “Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827610”; “IRAN / Keredj (1200 m.) / 4.4. 1936 / Brandt” // “Cotypus / P. ergane / ssp. pfeifferi / By.S & Br.” // “ ssp. pfeifferi Byt.” // “Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827611”; “IRAN / Keredj (1500 m.) / 20/5 1936 / Brandt” // “Cotypus / ergane ssp. / pfeifferi / By.S+Br.” // “Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827612”; “IRAN / Keredj (1500 m.) / 18/5 1936 / Brandt” // “Cotypus / ergane ssp. / pfeifferi / By.S+Br.” // “ssp. pfeifferi / By.S. & Br.” // “Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827613”.

Type locality

Iran: Tehran, “Keredj” [Karaj].

Remarks

Label indicates ssp. pfeifferi, but apparently this name does not exist. Collection data are identical to ssp. elbursina, possibly a last-minute name change by Bytinski-Salz.

r. exigua Verity, 1923

Original combination

Pieris ergane, H.-G. race exigua, mihi.” Verity, 1923 Ent. Rec. 35 Suppl. (18).

Current combination

Pieris ergane r. exigua Verity, 1923.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Cotype” 2♂♂ (ZMH 835577–835578) (Fig. 65). “exigua Vrty” // “Marche / Bolognola / 7 Ag 1937 / Querci” // [blank label] // “ZMH 835577”; “exigua Vrty” // “Marche / Bolognola / 8 Ag 1937 / Querci” // [blank label] // “ZMH 835578”.

Original locality

Italy: “Upper Fargno Valley, at 1400 m., in the Sibillini Mts.”

Remarks

Verity (1923) proposed this name as a race of P. ergane (Geyer, 1828). According to Kudrna (1983), this name in unavailable because it is a seasonal form and also a nomen nudum. The year on the specimen labels (1937) indicate specimens were collected 14 years after description (1923) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these two specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.

f. rossioides Stauder, 1921

Original combination

P. rapae L. f. n. rossioides” Stauder, 1921 Dt. Ent. Z. Iris 35: 27.

Current combination

Pieris rapae f. rossioides Stauder, 1921.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827627) (Fig. 66). “Calabria mer. / Aspromonte 800 m. / 4/7 [illegible] 1920 / H. Stauder legit.” // “Type” // “Rossioides Stdr.” // “rossioides / Stdr. / Type” // “ZMH 827627”.

Original locality

Italy: “Polsibeken, Aspromonte 7–900 m.”

Remarks

Stauder (1921) proposed this name as a form of P. rapae (Linnaeus, 1758). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work (please see below) unambiguously reveals that the name was proposed for an infrasubspecific entity. In the same paper, Stauder (1921) described another “form” of P. rapae as “zelleri” and clearly stated that it also occurs at the same “type locality” as rossioides (i.e., Aspromonte). As by definition two geographical races (or forms) cannot occur sympatrically; therefore, the content of the work reveals that such forms were not meant to be subspecific. However, Stauder added further complexity; there is a difference between how “zelleri” and “rossioides” have been described. In fact, the former was as ‘g. aest. mer. alt.’, i.e., generatio aestivalis meridionalis altitudinalis [Southern altitudinal summer generation], while the latter just as ‘f. n.’, i.e., “forma nova” [new form]. Thus, it could theoretically be argued that rossioides was meant as a geographical race, while zelleri would be a seasonal variation of that. However, this speculation does not seem to be correct, because it makes no sense that someone described a seasonal form of a subspecies before describing the subspecies itself. As a result, looking at the paper as a whole and not to the isolated description of rossioides alone, this name has to be considered as infrasubspecific (Alberto Zilli pers. comm.).

Unpublished, unavailable names

ab. gottingensis Meinhard

Original combination

Pieris napi v. napaeae ab. gottingensis Meinhard.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 835579) (Fig. 67). “v. gottingensis / Mht” // “8/5 1909 / Göttingen / Hainholz” // “ZMH 835579”.

Remarks

This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable.

ab. nigromarginata Meinhard

Original combination

Pieris brassicae ab. nigromarginata Meinhard.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 835576) (Fig. 68). “13/8 1916 / Göttingen / Hainberg” // “ZMH 835576”.

Remarks

This taxon cannot be found in the literature. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

Family Lycaenidae Leach, 1815

Subfamily Aphnaeinae Distant, 1884

dueldueli Pfeiffer, 1932

Original combination

Cigaritis acamas Klug. ssp. n. düldüli m.” Pfeiffer, in Osthelder and Pfeiffer 1932, Mitt. Münch. Ent. Ges. 22: 38.

Current combination

Cigaritis acamas dueldueli Pfeiffer, 1932.

Current status

Valid subspecies.

Type material

Syntypes 3♂♂ (ZMH 827553–827555) (Fig. 69). “Syria s. / amanus / Düldül Dagh / VII.30” // “Cotypus / A. epargyros / ssp. düldüli / Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827553”; “Syria sept. / amanus sept. / Düldül Dagh / VII.30” // “Cotypus / A. epargyros / ssp. düldüli / Pfeiffer” // “Coll. Bytinski Salz / Eing. Nr. 20, 1960” // “ZMH 827554”; “Syria sept. / amanus sept. / Düldül Dagh / VII.30” // “Cotypus / A. epargyros / ssp. düldüli / Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827555”.

Type locality

Turkey: “Marasch in türkisch Nordsyrien; Im DD [Düldül Dagh].”

Subfamily Lycaeninae Leach, 1815

Tribe Lycaenini Leach, 1815

f. emarginata Warnecke, 1942

Original combination

Chrysophanus alciphron Rott., ♂, n. f emarginata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.

Current combination

Lycaena alciphron f. emarginata (Warnecke, 1942).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827527) (Fig. 70). “Schlesw. – Holst. / Kr. / Bockhorn / W. Wolf leg. / 14.7.40” // “Katalog Nr. / L 2322” // “♂ nf. emarginata / Warn. Type / Iris 1942, 103” // “Chrysophanus / alciphron / emarginata Warn. / Type / L.31” // “Sig. G. Warnecke / Eing. Nr. 5, 1945” // “ZMH 827527”.

Original locality

Germany: “Segeberg”; Schleswig-Holstein publication says Segeberg; the label of the specimen treated as type by Warnecke states Lauenburg crossed out and replaced by Bockhorn.

Remarks

Warnecke (1942) proposed this name as a form of C. alciphron (Rottemburg, 1775). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. Already the title of Warnecke (1942) paper reveals that this form was described as an aberration and not as a subspecies.

ab. alba Schoenfeld, 1924

Original combination

Chrysophanus virgaureae L.-♀, alba m., nov. ab.” Schoenfeld, 1924 Int. Ent. Z. 18: 40.

Current combination

Lycaena virgaureae ab. alba (Schoenfeld, 1924).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♀ (ZMH 827586) (Fig. 71). “Bayern / Mitterteich / 24.VIII.1923 / Schönfeld” // [blank label] // “Type / Chr. virgaureae L. / ab. alba Schönfeld / I. E. Z. Guben 18. Jahrg / 1924. Nr 6. P. 40 / [illegible]” // “ZMH 827586”.

Original locality

Germany: Bavaria.

Remarks

Schoenfeld (1924) proposed this name as an aberration of C. virgaureae (Linnaeus, 1758), therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

f. multipunctata Warnecke, 1942

Original combination

Chrysophanus virgaureae L., ♂, n.f. multipunctata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.

Current combination

Lycaena virgaureae f. multipunctata (Warnecke, 1942).

Current status

Infrasubspecific and hence unavailable.

Original material

Labelled as “Type” 1♂ (ZMH 827592) (Fig. 72). “n.f. multipunctata / Warn. Type / Iris 1942, 103” // “Chrysoph. virgaureae / multipunctata / Type Warn. / L.30” // “Holstein” // “Katalog Nr. / L. 2196” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827592”.

Original locality

Germany: Holstein.

Remarks

Warnecke (1942) proposed this name as a form of C. virgaureae (Linnaeus, 1758). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. The title and the text of Warnecke (1942) paper unambiguously reveals that this form was described as an aberration and not as a subspecies. It was only found in a single male.

f. quercii Turati, 1923

Original combination

Chrysophanus virgaureae emilianus f. n. e. quercii f. n.” Turati, 1923 Atti. Soc. Ital. Sci. Nat. 62: 42.

Current combination

Lycaena virgaureae emilianus f. quercii (Turati, 1923).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 6♂♂ (ZMH 827580–827585) (Fig. 73). “Marche / Bolognola / 15.VII.1937 / Querci” // [blank label] // “quercii Trti.” // “ZMH 827580”; “Marche / Bolognola / 15.VII.1937 / Querci” // [blank label] // “quercii Trti.” // “ZMH 827581”; “Marche / Bolognola / 3 Ag 1937 / Querci” // [blank label] // “quercii Trti.” // “ZMH 827582”; “Marche / Bolognola / 27.VII.1937 / Querci” // [blank label] // “quercii Trti.” // “ZMH 827583”; “Marche / Bolognola / 5 Ag 1937 / Querci” // [blank label] // “quercii Trti.” // “ZMH 827584”; “Marche / Bolognola / 4 Ag 1937 / Querci” // [blank label] // “quercii Trti.” // “ZMH 827585”.

Original locality

Italy: “Monti Sibillini”.

Remarks

Turati (1923) proposed this name as a form of C. v. emilianus Turati, 1923. Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable.

Subfamily Polyommatinae Swainson, 1827

Tribe Polyommatini Swainson, 1827

andarabi Forster, 1938

Original combination

Lycaena pheretiades andarabi ssp. nov.” Forster, 1937 Mitt. Münch. Ent. Ges. 27: 61.

Current combination

Agriades pheretiades andarabi (Forster, 1938).

Current status

Valid subspecies.

Type material

Paratypes 4♂♂ (ZMH 827575–827578) (Fig. 74). “West-Hindukusch / Andarab / Geröllzone / 4000–4500 m Mitte August / leg. H.&E. Kotzsch” // “Para-Type / Lyc. Pheretiades / andarabi Forst. / W. Forster München” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827575”; “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. Pheretiades / andarabi Forest. / W. Forster München” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827576”; “West-Hindukusch / Andarab / Geröllzone / 4000–4500 m Mitte August / leg. H.&E. Kotzsch” // “Para-Type / Lyc. Pheretiades / andarabi Forest. / W. Forster München” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827577”; “West-Hindukusch / Andarab / Geröllzone / 4000–4500 m Mitte August / leg. H.&E. Kotzsch” // “Para-Type / Lyc. Pheretiades / andarabi Forest. / W. Forster München” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827578”.

Type locality

Afghanistan: “West-Hindukusch, Andarab, 4000–4500 m.”

f. altera Züllich, 1929

Original combination

Lycaena anteros Frr. f. altera” Züllich, 1929 Z. Öst. Ent. Ver. 14: 52.

Current combination

Aricia anteros f. altera (Züllich, 1929).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type”1♂ (ZMH 827526) (Fig. 75). “BULGARIA OR. / GIOKTEPE 250 M / MITTE AUG 1928 / COLL. ZUELLICH” // “ExTypicalibus / Specimenibus” // “VIII 1928 / Giöktepe, / Bulgarien” // “ZMH 827526”.

Original locality

Bulgaria: “Rilogebiet, Südwestbulgarien, 1200 m, und Gioktepe, Südostbulgarien, 200 m.”

Remarks

Züllich (1929) proposed this name as a form of L. anteros Freyer, 1839. According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. The text of Züllich (1929: 52) unambiguously reveals that this form was described explicitely for the summer generation.

r. modicior Verity, 1938

Original combination

Aricia anteros, Frr. modicior, nom. nov.” Verity, 1938 Ent. Rec. 50(8): Suppl. (5).

Current combination

Aricia anteros r. modicior Verity, 1938.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 2♂♂, (ZMH 827535–827536) (Fig. 76). “modicior Vrty” // “Macedonia / Olympus, 4500’ / July 2, 1935 / Romei” // [blank label] // “ZMH8274535”; “modicior Vrty” // “anteros Frr.” // “Macedonia / Olympus, 4500’ / July14, 1935 / Romei” // [blank label] // “ZMH8274536”.

Original locality

Greece: “At the highest altitudes of Mt. Olympus, such as at Prionia, 1500 m”.

Remarks

According to Kudrna (1983: 32), this name is unavailable and denotes a race and seasonal form and is hence unavailable.

uliginosa Dannehl, 1921

Original combination

Lycaena argus Schiff. var. uliginosa” Dannehl, 1921 Mitt. Münch. Ent. Ges. 11: 39.

Current combination

Plebejus idas uliginosa (Dannehl, 1921).

Current status

Junior subjective synonym of Plebejus argus argus (Linnaeus, 1758).

Type material

Syntypes 4♂♂, (ZMH 827541–827542, ZMH 835595–835596) (Fig. 77). “18.7” // “Ob. Bayern / Beuerberg / c. Dannehl 20” // “Argus v. n. / uliginosa” // “Katalog Nr. / L 2562” // “Cotype” // “L.34 / Lyc. argus - / v. uliginosa Dannehl / ♂ Cotype” // Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827541”; “18.7” // “Ob. Bayern / Beuerberg / c. Dannehl 20” // “v. uliginosa / Dannehl” // “Cotype” // “Katalog Nr. / L 2561” // “L.35 / Lyc. argus - / v. uliginosa Dannehl / ♀ Cotype” // Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827542”; “27.6” // Ob. Bayern / Beuerberg / F. Dannehl” // “Cotype” // “27.6. 1921 / Beuerberg / Oberbayern / v. Dannehl” // “ZMH 835595”; “13.7” // Ob. Bayern / Beuerberg / F. Dannehl” // “Cotype” // “13.7. 1921 / Beuerberg / Oberbayern / v. Dannehl” // “ZMH 835596”.

Type locality

Germany: Bavaria, Beuerberg.

Remarks

Dannehl (1921) proposed this name as a “var.” of L. argus ([Denis & Schiffermüller], 1775). L. argus is a junior subjective synonym of Plebejus idas Linnaeus, [1760]. According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.

balcanica Züllich, 1929

Original combination

Lycaena argyrognomon Bgstr. f. balcanica” Züllich, 1929 Z. Öst. Ent. Ver. 14: 52.

Current combination

Plebejus argyrognomon balcanica (Züllich, 1929).

Current status

Preoccupied by Lycaena semiargus balcanica Tutt, 1909; replacement name Lycaeides argyrognomon baldur (Hemming, 1934) is the current valid name.

Type material

Holotype 1♂ (ZMH 827544) (Fig. 78) paratype 1♂ (ZMH 827543). “BULGARIA OCC. // MON. RILSKI 1200 M / 20. – 31 VII. 1928 // COLL. ZUELLICH” // “Ex Typicalibus / Speciminibus” // “Holotype” // “Coll. Bytinski-Salz / Eing Nr 20, 1960” // ZMH 827544”; “BULGARIA OCC. // MON. RILSKI 1400 M / 20 VI. – 6 VII. 1928 // COLL. ZUELLICH” // “Ex Typicalibus / Speciminibus” // “Paratypoid” // “Coll. Bytinski-Salz / Eing Nr 20, 1960” // ZMH 827543”.

Type locality

Bulgaria: “MON. RILSKI” [Monastery Rilski].

Remarks

Züllich (1929) proposed this name as a form of L. argyrognomon (Bergsträsser, 1779). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “form”.

mayencis Eitschberger & Steiniger, 1975

Original combination

Eumedonia eumedon mayencis n. subsp.” Eitschberger & Steiniger, 1975 Atalanta 6 (1): 106.

Current combination

Eumedonia eumedon mayencis Eitschberger & Steiniger, 1975.

Current status

Valid subspecies.

Type material

Paratypes 5♂♂ (ZMH 827548–827552) (Fig. 79). “Gonsenheim / 5. Juni / 04 / ANDREAS / Gonsenheim” // “PARATYPUS o / Eumedonia eu- / medon mayencis / EITSCHBERGER & / STEINIGER, 1975” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827548”; “Gonsenheim / 5. Juni / 04 / ANDREAS / Gonsenheim” // “PARATYPUS o / Eumedonia eu- / medon mayencis / EITSCHBERGER & / STEINIGER, 1975” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827549”; “Mainz / Mombach / 10.6.1922” // “Eumedon” // “PARATYPUS o / Eumedonia eu- / medon mayencis / EITSCHBERGER & / STEINIGER, 1975” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827550”; “Gonsenheim / 8. Juni / 04 / ANDREAS / Gonsenheim” // “PARATYPUS o / Eumedonia eu- / medon mayencis / EITSCHBERGER & / STEINIGER, 1975” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827551”; “Mainz / Mombach / 10.6.1922” // “Eumedon” // “PARATYPUS o / Eumedonia eu- / medon mayencis / EITSCHBERGER & / STEINIGER, 1975” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827552”.

Type locality

Germany: “Deutschland, Rheinland-Pfalz, Mainz-Mombach, Mainzer Sand”.

moenus Eitschberger & Steiniger, 1975

Original combination

Eumedonia eumedon moenus n. subsp.” Eitschberger & Steiniger, 1975 Atalanta 6 (2): 99.

Current combination

Eumedonia eumedon moenus Eitschberger & Steiniger, 1975.

Current status

Valid subspecies.

Type material

Paratypes 2♂♂ (ZMH 827546–827547) (Fig. 80). “Tiertalsberg bei / Retzbach am Main / 4.6.1950” // “Katalog Nr. / L 3370” // “PARATYPUS o / Eumedonia eume- / don moenus / EITSCHBERGER & / STEINIGER, 1975” // “Bot.-Zool. Excursion / 1950/ Eing Nr. 14—1950” // “ZMH 827546”; “Karlstadt a. Main / Karlsburg / 4.6.1950” // “Katalog Nr. / L 3369” // “PARATYPUS o / Eumedonia eume- / don moenus / EITSCHBERGER & / STEINIGER, 1975” // “Bot.-Zool. Excursion / 1950/ Eing Nr. 14—1950” // “ZMH 827547”.

Type locality

Germany: “Deutschland, Nordbayern, mittleres Maintal und Nord-Baden, mittleres Taubertal.”

persa Bytinski-Salz & Brandt, 1937

Original combination

Chilades trochylus, Frr. ssp. persa, ssp. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 49: Suppl. (2).

Current combination

Freyeria trochylus persa (Bytinski-Salz & Brandt, 1937).

Current status

Valid subspecies.

Type material

Syntypes 4♂♂ (ZMH 827587–827590) (Fig. 81). “IRAN / Keredj (1600 m.) / 3/6 1936 / Brandt” // “Cotypus / Ch. trochylus / ssp. persa / By.S + Br.” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827587”; “IRAN / Keredj (1600 m.) / 9/8 1936 / Brandt” // “Cotypus / Ch. trochylus / ssp. persa / By.S + Brandt” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827588”; “IRAN / Keredj (1600 m.) / 12/8 1936 / Brandt” // “Cotypus / Ch. trochylus / ssp. persa / By.S + Br.” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827589”; “IRAN / Keredj (1600 m.) / 3/6 1936 / Brandt” // “Cotypus / Ch. trochylus / ssp. persa / ByS + Br.” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827590”.

Type locality

Iran: Tehran, “Keredj” [Karaj].

ab. pauper Bytinski-Salz & Brandt, 1937

Original combination

Chilades trochylus, Frr. ssp. persa, ssp. nov. ab. pauper, ab. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 49: Suppl. (2).

Current combination

Freyeria trochylus persa ab. pauper (Bytinski-Salz & Brandt, 1937).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Holotype” ♀ (ZMH 827591) (Fig. 82). “IRAN / Keredj (1600 m.) / 25/5 1936 / Brandt” // “ab. pauper / ByS+Br. / Holotype ♀ / ex. coll. / Dr. H. Bytinski-Salz” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827591”.

Original locality

Iran: Tehran, “Keredj” [Karaj].

Remarks

Bytinski-Salz and Brandt (1937) proposed this name as an aberration of C. trochylus (Freyer, 1838). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

parvandereggi Verity, 1938

Original combination

Glaucopsyche cyllarus, Rott, race parvandereggi, nom. nov.” Verity, 1938 Ent. Rec. 50: Suppl. (4).

Current combination

Glaucopsyche alexis parvandereggi Verity, 1938.

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 833470–833471) (Fig. 83). “parvandereggi Vrty” // “Macedonia / Olympus, 2900’ / V. 25, 1936 / Romei” // [blank label] // “ZMH 833470”; “parvandereggi Vrty” // “Macedonia / Olympus, 2900’ / V. 26, 1936 / Romei” // [blank label] // “ZMH 833471”.

Type locality

Greece: Salonika [Macedonia, Olympus].

Remarks

Verity (1938) proposed this name as a race of G. cyllarus (Rottemburg, 1775). As it was given in trinomy and the content of the description clearly indicates that a geographical race was meant, this name is fully available as subspecific (article 45.6 ICZN 1999). (Alberto Zilli pers. comm.).

sibyllina Verity, 1916

Original combination

Lycaena coridon” Poda, “razza sibyllina” Verity, 1916 Boll. Soc. Ent. Ital. 46: 133.

Current combination

Lysandra coridon sibyllina (Verity, 1916).

Current status

Junior subjective synonym of Lysandra coridon apennina (Zeller, 1847).

Original material

Labelled as “Type” 4♂♂ (ZMH 833493–833496) (Fig. 84). “sibyllina Vrty” // “Marche / Bolognola / 15 Ag 1937 / Querci” // [blank label] // “ZMH 833493”; “sibyllina Vrty” // “Marche / Bolognola / 9 Ag 1937 / Querci” // [blank label] // “ZMH 833494”; “sibyllina Vrty” // “Marche / Bolognola / 9 Ag 1937 / Querci” // [blank label] // “ZMH 833495”; “sibyllina Vrty” // “Marche / Bolognola / 8 Ag 1937 / Querci” // [blank label] // “ZMH 833496”.

Original locality

Italy: “Monti Sibillini (Bolognola, 1200 m)”

Remarks

Verity (1938) proposed this name as a “razza” (=race) of L. coridon (Poda, 1761). As it was given in trinomy and the content of the description clearly indicates that a geographical race was meant, this name is available as subspecific (article 45.6 ICZN 1999). However, it was later synonimized with L. coridon apennina (Zeller, 1847). The year on the specimen labels (1937) indicate specimens were collected 21 years after description (1916) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these four specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.

f. lunulata Warnecke, 1942

Original combination

Lycaena alcon F., ♀, n. f. lunulata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.

Current combination

Phengaris alcon f. lunulata (Warnecke, 1942).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♀ (ZMH 827528) (Fig. 85). “Bahrenfeld / 28.7.08” // “Sig. S. Warnecke / Eing. Nr. 5 1949” // “♀ n.f. lunuata / Warn. Type / Iris 1942, 103” // “Katalog Nr. L 32 67” // “ZMH 827528”.

Original locality

Germany: Eidelstedt, Hamburg.

Remarks

Warnecke (1942) proposed this name as a form of “L. alcon F.” (sic, recte: [Denis & Schiffermüller], 1775). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. The title of the paper (Warnecke, 1942) indicates that this form constitutes an aberration and is therefore infrasubspecific and unavailable. According to the description this form was only found in a single female.

selzeri Warnecke, 1924

Original combination

Lycaena arionides Staudinger f. selzeri” Warnecke, 1924 Dt. Ent. Z. Iris 38: 149.

Current combination

Phengaris arionides f. selzeri (Warnecke, 1924).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827545) (Fig. 86). “Ussuri” // “F. selzeri Warn. / ♂ Type” // “n.f. selzeri Warn. / I. Iris 1924, 38. J. / p. 149, Taf. II bis. 5.” // “Type” // “Lyc. arionides - / n.f. selzeri Warn. / L.33” // “Katalog Nr. / L 3336” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827545”.

Original locality

Russia: “Ostsibirien, Amur-Gebiet, Ussuri”.

Remarks

Warnecke (1942) proposed this name as a form of L. arionides Staudinger, 1887. According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. According to the description this form is an individual aberration or a rare second generation and is therefore infrasubspecific. Accordingly, on plate II this form is figured as “Lycaena arionides nov. ab. selzeri Warn.”.

peninsulae Verity, 1923

Original combination

Lycaena euphemus, Hüb. (a) race peninsulae, mihi” Verity, 1923 Ent. Rec. 35 Suppl. (7).

Current combination

Phengaris euphemus peninsulae (Verity, 1923).

Current status

Valid subspecies.

Original material

Labelled as “Type” 1♂ (ZMH 833468) (Fig. 87). “peninsulae Vrty” // “Marche / Bolognola / 10.VI.1937 / Querci / 1300 m” // [blank label] // “ZMH 833468”.

Original locality

Italy: “Sibillini Mts. in the Marche, below Bolognola, at 1000 m”.

Remarks

Verity (1923) proposed this name as a race of L. euphemus Hübner, 1799–1800. According to Kudrna (1983: 35), this race represents an available subspecific name. However, the year on the specimen labels (1937) indicate specimens were collected 14 years after description (1923) from the same location (i.e., Italy: Bolognola) the original type material was collected. As a result, the black label of the specimens treated as “type” by Verity is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, this specimen is erroneously labelled as type, should be annotated as “Not a Type” and are treated here as “Non-type” specimen.

ab. depuncta Hirschke, 1903

Original combination

Lycaena Argiades ab. (et var.) depuncta” Hirschke, 1903 Verh. Ges. Wien. 53: 270.

Current combination

Cupido argiades ab. depuncta (Hirschke, 1903).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827531) (Fig. 88). “Hirschke.Wien / Herkulsbad / V.1900 Type” // “Verh. zool. / bot. ges. / 1903 Hirs.” // “TYPE” // “ZMH 827531”.

Original locality

Romania: “Herkulesbad”.

Remarks

Hirschke (1903) proposed this name as an aberration of L. argiades (Pallas, 1771). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

ausonia Verity, 1913

Original combination

Lycaena damon ausonia nom. nov.” Verity, 1913 Boll. Soc. Ent. Ital. 45: 232, pl. 1, f. 44, 47.

Current combination

Polyommatus (Agrodiaetus) damon ausonia (Verity, 1913).

Current status

Junior subjective synonym of Polyommatus (Agrodiaetus) damon damon ([Denis & Schiffermüller], 1775).

Original material

Labelled as “Type” 6♂♂ (ZMH 833479–833484) (Fig. 89). “ausonia Vrty.” // “Marche / Bolognola / 15.VII.1937 / Querci” // [blank label] // “ZMH 833479”; “ausonia Vrty.” // “Marche / Bolognola / 15.VII.1937 / Querci” // [blank label] // “ZMH 833480”; “ausonia Vrty.” // “Marche / Bolognola / 12.VII.1937 / Querci” // [blank label] // “ZMH 833481”; “ausonia Vrty.” // “Marche / Bolognola / 5 Ag 1937 / Querci” // [blank label] // “ZMH 833482”; “ausonia Vrty.” // “Marche / Bolognola / 3 Ag 1937 / Querci” // [blank label] // “ZMH 833483”; “ausonia Vrty.” // “Marche / Bolognola / 12.VII.1937 / Querci” // [blank label] // “ZMH 833484”.

Original locality

Italy: “a grandi altitudini nei soliti monti del Piceno” [At high elevations at Mount del Piceno].

Remarks

The year on the specimen labels (1937) indicate specimens were collected 24 years after description (1913) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these six specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.

pfeifferi Brandt, 1938

Original combination

Lycaena Pfeifferi sp. n.” Brandt, 1938 Ent. Rdsch. 55: 671, pl. 4, f. 1–3.

Current combination

Polyommatus (Agrodiaetus) pfeifferi (Brandt, 1938).

Current status

Valid species.

Type material

Paratype 1♂ (ZMH 827579) (Fig. 90). “IRAN. FARS, Straße / Ardekan – Talochosroe / Comé, m, 1937 / coll. Brandt” // “L.37 / Paratypoid / Lyc. pfeifferi Brand” // “Katalog Nr. / L. 3190” // “Sammlung R. Jähnig / Eing. Nr.29, 1949” // “ZMH 827579”.

Type locality

Iran: Fars “vom Barm I Firus (3750 m)” [from Barm-e Firuz].

alcestis Zerny, 1932

Original combination

Lycaena (Hirsutina) ripperti alcestis Zerny nov. subsp.” Zerny, 1932 Dt. Ent. Z. Iris 46: 186.

Current combination

Polyommatus (Agrodiaetus) alcestis (Zerny, 1932).

Current status

Valid species.

Type material

Syntypes 2♂♂ (ZMH 827529–827530) (Fig. 91). “Syria / Libanon s. / Bscharre / VIII-IX 31 / 1300 m / Einh.Slr.leg” // “Cotypus / ripperti / ssp. alcestis / Zerny” // “Coll. Bytinski-Salz / Eing Nr 20, 1960” // “ZMH 827529”; “Syria / Libanon s. / Bscharre / VIII-IX 31 / 1300 m / Einh.Slr.leg” // “Cotypus / ripperti / ssp. alcestis / Zerny” // “Coll. Bytinski-Salz / Eing Nr 20, 1960” // “ZMH 827530”.

Type locality

“Lebanon, Becharré, 1400 m; Zedernwald oberhalb Becharré um 1900 m”.

ardschira Brandt, 1938

Original combination

Lycaena ardschira sp. n.” Brandt, 1938 Ent. Rdsch. 55: 672.

Current combination

Polyommatus (Agrodiaetus) ardschira (Brandt, 1938).

Current status

Valid species.

Type material

Paratype 1♂, (ZMH 827532) (Fig. 92). “L.36 / Paratypoid / Lyc. ardschira / Brand” // “Paratypoid” // “IRAN. FARS, Straße / Ardekan – Talochosroe / Comé, m, 1937 / coll. Brandt” // “196” // “Sammlung / R. Jähnig / Eing. Nr. 29,1949” // “Katalog Nr. / L. 3365” // “ZMH 827532”.

Type locality

Iran: Fars, “Comée, 2600 m” [Komehr].

f. squalida Verity, 1920

Original combination

Polyommatus meleager, Esp., race macra, mihi, ♀ form squalida, mihi” Verity, 1920 Ent. Rec. 32:145.

Current combination

Polyommatus daphnis macra f. squalida Verity, 1920.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 2♂♂ (ZMH 833485–833486) (Fig. 93). “squalida Vrty” // “Marche / Bolognola / 15.VII.1937 / Querci” // [blank label] // “ZMH 833485”; “squalida Vrty” // “Marche / Bolognola / 3 Ag 1937 / Querci” // [blank label] // “ZMH 833486”.

Original locality

Italy: “Mt. Sibillina, 1200–1300 m”.

Remarks

Verity (1920) proposed this name as a form of P. meleager macra Verity, 1920. According to Kudrna (1983: 39), this name is unavailable and denotes a race and is infrasubspecific and hence unavailable. Papilio meleager Esper, 1779 is a junior subjective synonym of Papilio daphnis [Denis & Schiffermüller], 1775. The year on the specimen labels (1937) indicate specimens were collected 17 years after description (1920) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these two specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.

r. correpta Verity, 1920

Original combination

Agriades hylas, Esp., race correpta, mihi” Verity, 1920 Ent. Rec. 32: 144.

Current combination

Polyommatus dorylas r. correpta (Verity, 1920).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 4♂♂ (ZMH 833489–833492) (Fig. 94). “correpta Vrty” // “Marche / Bolognola / 15.VII.1937 / Querci / 1500 m” // [blank label] // “ZMH 833489”; “correpta Vrty” // “Marche / Bolognola / 14.VII.1937 / Querci / 1400 m” // [blank label] // “ZMH 833490”; “correpta Vrty” // “Marche / Bolognola / 29.VII.1937 / Querci / 1600 m” // [blank label] // “ZMH 833491”; “correpta Vrty” // “Marche / Bolognola / 12.VII.1937 / Querci / 1500 m” // [blank label] // “ZMH 833492”.

Original locality

Italy: “Bolognola, Central Italy”.

Remarks

Verity (1920) proposed this name as a race of A. hylas (Esper, 1779). According to Kudrna (1983: 26), this name is unavailable as it denotes a seasonal form which is infrasubspecific and hence unavailable. The year on the specimen labels (1937) indicate specimens were collected 17 years after description (1920) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these four specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.

olympena Verity, 1936

Original combination

L. escheri, Hüb. race olympena, nom. nov.” Verity, 1936 Ent. Rec. 48: 8.

Current combination

Polyommatus escheri olympena (Verity, 1936).

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 833487–833488) (Fig. 95). “olympena Vrty” // “Macedonia / Olympus, 2500’ / July 13, 1935 / Romei” // [blank label] // “ZMH 833487”; “olympena Vrty” // “Macedonia / Olympus, 2500’ / July 9, 1936 / Romei” // [blank label] // “ZMH 833488”.

Type locality

Greece: “Mt. Olympus, from 850 to 1500 m.”

Remarks

Verity (1936) proposed this name as a race of L. escheri Hübner, 1823. According to Kudrna (1983: 34), this race represents an available subspecific name.

ab. privata Schönfeld, 1924

Original combination

Lycaena icarus” Rott.-♂ “ab. privata m., nov. ab.” Schoenfeld, 1924 Int. Ent. Z. 18: 40.

Current combination

Polyommatus icarus ab. privata (Schönfeld, 1924).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827573) (Fig. 96). “14.5. 1921 / Kl. Chuchel / Böhmen / von Schönfeld” // “Type / Lyc. icarus Rott. ♂ / ab. privata Schönfeld / I.E.Z. Guben. 18. Jahr. Nr 6 / 1924. a.d. nachl. v. Schönfeld. / D. Gelpke” // “Kl – Chuchel / 14.5.21” / “ZMH 827573”.

Original locality

Czech Republic: “in der Umgebung von Prag (bei Kuchelbad)“ [in the vicinity of Prague (near Kuchelbad)].

Remarks

Schoenfeld (1924) proposed this name as an aberration of L. icarus Rottemburg, 1775. Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

nuksani Forster, 1937

Original combination

Lycaena candalus nuksani ssp. nov.” Forster, 1937 Mitt. Münch. Ent. Ges. 27: 62.

Current combination

Polyommatus (Sublysandra) nuksani (Forster, 1937).

Current status

Valid species.

Type material

Paratypes 2♂♂ 2♀♀, (ZMH 827537–827540) (Fig. 97). “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. candalus / nuksani Forst. / W. Forster München” // “Coll. Bytinski-Salz / Eing. Nr. 20 – 60” // “ZMH 827537”; “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. candalus / nuksani Forst. / W. Forster München” // “Coll. Bytinski-Salz / Eing. Nr. 20 – 60” // “ZMH 827538”; “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. candalus / nuksani Forst. / W. Forster München” // “candalus / nuksani” // “Coll. Bytinski-Salz / Eing. Nr. 20 – 60” // “ZMH 827539”; “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. candalus / nuksani Forst. / W. Forster München” // “Coll. Bytinski-Salz / Eing. Nr. 20 – 60” // “ZMH 827540”.

Type locality

Afghanistan: “Nordost-Hindukusch, Nuksan-Paß Nordseite, 3500–4000 m.”.

clorinda Verity, 1938

Original combination

Tarucus balkanica, Freyer, with I gen. clorinda, nom. nov.” Verity, 1938 Ent. Rec. 50: (8).

Current combination

Tarucus balkanica ab. clorinda Verity, 1938.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 2♂♂ (ZMH 835592–835593) (Fig. 98). “clorinda Vrty” // “Macedonia / Salonika, 1000’ / May 31, 1935 / Romei” // [blank label] // “ZMH 835592”; “clorinda Vrty” // “Macedonia / Salonika, 1000’ / May 31, 1935 / Romei” // [blank label] // “ZMH 835593”.

Original locality

Greece: “Salonika”.

Remarks

Verity (1938) proposed this name as an aberration of T. balkanica (Freyer, 1844). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

ahasveros Bytinski-Salz & Brandt, 1937

Original combination

Lycaena (Glaucopsyche) panagaea, H.-Sch. ssp. ahasveros ssp. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 49: Suppl. (2).

Current combination

Turanana panagaea ahasveros (Bytinski-Salz & Brandt, 1937).

Type material

Paratypes 14♂♀ (ZMH 827556–827569) (Fig. 99). “IRAN: / Keredj (1700 m.) / 6.5.1936 / Brandt” // “Cotypus / L. panagaea / ssp. ahasveros / ByS+Brdt” // “ ssp. ahasveros / ByS + Brdt.” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827556”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827557”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827558”; “IRAN: / Keredj (1200 m.) / 6.5.1936 / Brandt” // “Cotypus / L. panagaea / ssp. ahasveros / ByS+Brdt” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827559”; “IRAN: / Keredj (1700 m.) / 6.5.1936 / Brandt” // “Cotypus / L. panagaea / ssp. ahasveros / ByS+Brdt” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827560”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827561”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827562”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827563”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827564”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827565”; “IRAN: / Keredj (1700 m.) / 10.5.1936 / Brandt” // “Cotypus / L. panagaea / ssp. ahasveros / ByS+Brdt” // “ ssp. ahasveros / ByS + Brdt.” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827566”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827567”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827568”; “X” // “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827569”.

Type locality

Iran: Tehran, “Keredj 1700 m.” [Karaj].

Current status

Valid subspecies.

Subfamily Theclinae Swainson, 1831

Tribe Eumaeini Doubleday, 1847

syra Pfeiffer, 1932

Original combination

Thecla ilicis Esp. ssp. n. syra m.” Pfeiffer, 1932 in Osthelder and Pfeiffer 1932, Mitt. Münch. Ent. Ges. 22: 31.

Current combination

Satyrium ilicis syra (Pfeiffer, 1932).

Current status

Junior subjective synonym of nominotypical Satyrium ilicis (Esper, 1779).

Type material

Syntypes 3♂♂ (ZMH 827570–827572) (Fig. 100). “Syria sept. / amanüs / VI.1922” // “Cotypus / ilicis / ssp. syra / Pfeiffer” // “ssp. syra / Pfeiffer” // “Coll. Bytinski-Salz Coll. / Eing. Nr. 20, 1960 Eing.” // “ZMH 827570”; “Syria sept. / Taurus / Marasch / 26.V.28 / L. Osthelder leg. / 900 m” // “Cotypus / ilicis / ssp. syra / Pfeiffer” // “Coll. Bytinski-Salz Coll. / Eing. Nr. 20, 1960 Eing.” // “ZMH 827571”; “Syria sept. / Taurus / Marasch / 26.V.28 / L. Osthelder leg. / 1200 m” // “Cotypus / ilicis / ssp. syra / Pfeiffer” // “Coll. Bytinski-Salz Coll. / Eing. Nr. 20, 1960 Eing.” // “ZMH 827572”.

Type locality

Turkey: “Syria sept., Taurus, Marasch” [Marash].

brevicaudis Vorbrodt, 1911

Original combination

Thecla linceus Esp. (= spini Schiff.) brevicaudis Püngeler, i.l.” Vorbrodt, 1911 Die Schmetterlinge der Schweiz. I.: 106.

Current combination

Satyrium spini brevicaudis (Vorbrodt, 1911).

Current status

Junior subjective synonym of nominotypical Satyrium spini ([Denis & Schiffermüller], 1775).

Type material

Syntype 1♀ (ZMH 827574) (Fig. 101). “Zermatt / Rpe an Rhamnus alp.[ina] / myrmecophil mit / Formica fusca 6.1908 / ♀ c.l. 28.7.1908 Püng.” // “Cotype / v. brevicaudis Püng.” // “Spini var. brevicaudis Püng” // “L.29” // “Katalog Nr. / L 2048” // Sig. G. “Warnecke / Eing. Nr. 5, 1949” // “ZMH 827574”.

Type locality

Switzerland: “bei Zermatt 1700 m”.

Remarks

Vorbrodt (1911) proposed this name as a form (in the original description and “var.” on the specimen labels) of “T. Linceus” (sic, recte: lynceus) (Esper, 1779). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “form” or “var”.

Unpublished, unavailable names

v. romana Hartig

Original combination

Lycaena cyllarus v. romana Hartig.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Cotype” 3♂♂ (ZMH 833472–833474) (Fig. 102). “romana Hartig” // Lazio / Formia / 23 4 [19]38 / Querci” // [blank label] // “ZMH 833472”; “romana Hartig” // Lazio / Formia / 8 4 [19]38 / Querci” // [blank label] // “ZMH 833473”; “romana Hartig” // Lazio / Formia / 3 5 [19]38 / Querci” // [blank label] // “ZMH 833474”.

Original locality

Italy: Lazio.

Remarks

This taxon cannot be found in the literature. However, it may refer to Glaucopsyche alexis latina Hartig, 1940 (Mem. Soc. Ent. Ital. 18: 187; Type locality: Formia, Latina, Italy). The label data matches the type locality of latina. Papilio cyllarus Rottemburg, 1775 is a junior subjective synonym of Papilio alexis Poda, 1761.

ab. albipunctata Hartig

Original combination

Lysandra coridon ab. albipunctata Hartig.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 4♂♂ (ZMH 833475–833478) (Fig. 103). “Typen” // “19/7 1917 / Göttingen / Hohefeld” // “ZMH 833475”; “13/7 1917 / Göttingen / Hohefeld” // “ZMH833476”; “18/8 1917 / Göttingen / Hohefeld” // “ZMH833477”; “18/7 1917 / Göttingen / Hohefeld” // “ZMH 833478”.

Original locality

Germany: Göttingen.

Remarks

This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

Family Nymphalidae Rafinesque, 1815

Subfamily Danainae Boisduval, 1833

icensis Fuchs, 1954

Original combination

Danais gilippus var. icensis” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84.

Current combination

Danaus gilippus icensis (Fuchs, 1954).

Current status

Junior subjective synonym of Danaus eresimus erginus (Godman & Salvin, 1897).

Type material

Lectotype 1♂ (ZMH 827645), paralectotypes 2♂♂ (ZMH 827643–827644) (Fig. 104). “Huayuri / 5/1940 / Type” / “Coll. H. Rödinger / Eing. Nr. 30, 1952 / Katalog Nr. L” // “ZMH 827643”; “Ica / 7/1940 / Type” / “Coll. H. Rödinger / Eing. Nr. 30, 1952 / Katalog Nr. L” // “Paralectotype ♀ / Danais gilippus / var. icensis / von Fuchs / G. Lamas ‘98” // “Danaus eresimus / erginus Godman + / Salvin ♀ / G. Lamas. Det. 1978” // “ZMH 827644”; “114 / eresimus. ab / Ica (Perú)” // “Ica / 7.1928 / Type” // “Coll. H. Rödinger / Eing. Nr. 30, 1952 / Katalog Nr. L” // “Lectotype ♂ / Danais gilippus / var. icensis / von Fuchs / G. Lamas ‘78” // “Danaus eresimus / erginus Godman + / Salvin ♂ / G. Lamas. Det. 1978” // “ZMH 827645”.

Type locality

Peru: “Im Tal von Ica und des Río Grande”.

Remarks

Fuchs (1954) proposed this name as a “var.” of Danais gilippus Cramer, 1775. According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.

roedingeri Fuchs, 1954

Original combination

Danais gilippus var. Rödingeri” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84.

Current combination

Danaus gilippus roedingeri (Fuchs, 1954).

Current status

Junior subjective synonym of D. g. candidus Clark, 1941.

Type material

Holotype 1♂ (ZMH 827652), paratypes 6♂♀ (ZMH 827649–827651, ZMH 827653–827655) (Fig. 105). “Danais / gilippus / var. rödingeri ♂ / v. Fuchs” // “Type” // “Sammlung / W. v. Fuchs / Eing. Nr. 4, 1956” // “Süd – Peru / Ayacucho – Yucaes / 2500 m 26. 4. 1936.” // “Holotype ♂ / Danais gilippus / var. Rödingeri / von Fuchs” // “Danais gilippus ♂ / candidus Clark / G. Lamas. det. 1978” // “Hbg. – Süd – Peru / Sammelreise 1936. / Eing. Nr. 1, 1937.” // “ZMH 827652”; “Ayacucho / 2–3000 m / 8/1920 / Type” // “D. rödingeri ♀” // “Coll. H. Rödinger / Eing. Nr. 30, 1952” // “ZMH 827649”; “Type” // “Danais / gilippus // var. rödingeri ♂ / v. Fuchs / Rio Jucaes b/ / Ayacucho / 1800–2300” // “Summlung / W. v. Fuchs / Eing. Nr. 4, 1956” // “ZMH 827650”; “Ayacucho / 2–3000 m / 8/1920 / Type” // “Coll. H. Rödinger / Eing. Nr. 30, 1952 / Katalog Nr. L” // “ZMH 827651”; “Ayacucho 28.4” // “Süd – Peru / Oberh. Ayacucho / 29.4.1936.” // “Ayacucho / 2–3000 m / 8/1920 / Type” // “Summlung / W. v. Fuchs / Eing. Nr. 4, 1956” // “Hbg. – Süd – Peru / Sammelreise 1936. / Eing. Nr. 1, 1937.” // “ZMH 827653”; “Danais / gilippus / var. rödingeri ♀ / v. Fuchs” // “Type” // “Sammlung / W. v. Fuchs / Eing. Nr. 4, 1956” // “Süd – Peru / Ayacucho – Yucaes / 2500 m 26. 4. 1936.” // “Hbg. – Süd – Peru / Sammelreise 1936. / Eing. Nr. 1, 1937.” // “Danais gilippus ♀ / candidus Clark / G. Lamas. det. 1978” // “Paratype / Danaus gilippus ♀ / var. Rödingeri / von Fuchs” // “ZMH 827654”; “Type” // Danais / gilippus / var. rödingeri ♀ / v. Fuchs / Rio Jucaes / 6 / Ayacucho / 1800–2300” // “ZMH 827655”.

Type locality

Peru: “… näheren Umgebung von Ayacucho, auf 2000–2600 m” [vicinity of Ayacucho, 2000–2600 m].

Remarks

Fuchs (1954) proposed this name as a “var.” of Danais gilippus (Cramer, 1775). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.

Subfamily Heliconiinae Swainson, 1822

altonevadensis Reisser, 1927

Original combination

Argynnis niobe var. nov. altonevadensis” Reisser, 1927 Int. Ent. Z. Guben 20: 373.

Current combination

Fabriciana niobe altonevadensis (Reisser, 1927).

Current status

Junior subjective synonym of Fabriciana niobe niobe (Linnaeus, 1758).

Type material

Syntypes 1 ♂ 1♀ (ZMH 827656–827657) (Fig. 106). “Type # 7 ♂ / Argynnis niobe var. / altonevadensis Reisser / Int. Ent. Z. guben / Summer 1927” // “HISPANIA VII 26 / SIERRA NEVADA / O BUBACEK” // [blank label] // “ZMH 827656”; “Type # 4 ♀ / Argynnis niobe var. / altonevadensis Reisser / Int. Ent. Z. guben / Summer 1927” // “HISPANIA VII 26 / SIERRA NEVADA / O BUBACEK” // [blank label] // “ZMH 827657”.

Type locality

Spain: “… auf den Höhen der Sierra Nevada in Südspanien …“. [higher areas of Sierra Nevada in southern Spain].

Remarks

Reisser (1927) proposed this name as a “var.” of Argynnia niobe (Linnaeus, 1758). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.

cyrnea Schawerda, 1926

Original combination

Argynnis pandora Schiff. var. nov. cyrnea” Schawerda, 1926 Z. Öst. Ent. Ver. 11: 109.

Current combination

Argynnis pandora cyrnea Schawerda, 1926.

Current status

Junior subjective synonym of Argynnis pandora pandora ([Denis & Schiffermüller], 1775).

Type material

Syntypes 2♂♂ (ZMH 833447–833448) (Fig. 107). “pandora / var. cyrnea Schawerda / Cotype [illegible]” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 833447”; “pandora / var. cyrnea Schawerda / Cotype [illegible]” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 833448”.

Type locality

France: “Korsika” [Corsica].

Remarks

Schawerda (1926) proposed this name as a “var.” of Argynnis pandora ([Denis & Schiffermüller], 1775). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.

argentifasciata Kotzsch, 1937

Original combination

Argynnis pandora Schiff. argentifasciata ssp. n.” Kotzsch, 1937 Ent. Rdsch. 55: 9.

Current combination

Argynnis pandora argentifasciata Kotzsch, 1937.

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 827665–827666) (Fig. 108). “West-Hindukusch / Andarab / Geröllzone / 4000–4500 m Mitte August / leg. H.&E. Kotzsch” // “Cotypus / A. pandora / ssp. argenti- / fasciata / Kotzsch” // “ssp. argentifasciata / Kotzsch” // “ZMH827665”; “West-Hindukusch / Andarab / Banu-Ebene / 2000–2500 m August / leg. H.&E. Kotzsch” // “Cotypus / A. pandora / ssp. argenti- / fasciata / Kotzsch” // “ZMH827666”.

Type locality

Afghanistan: “Andarab im westlichen Hindukusch”.

Subfamily Nymphalinae Rafinesque, 1815

Tribe Melitaeini Newman, 1870

f. urbanoides Warnecke, 1942

Original combination

Melitaea maturna L., ♂, n. f. urbanoides.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.

Current combination

Euphydryas maturna f. urbanoides (Warnecke, 1942).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 2♂♂ (ZMH 827658–827659) (Fig. 109). “Holstein / Innien, e.l. / 1913 Selzer” // “♂ nf. urbanoides / Warn. Type / Iris 1942, 103” // “L.25 / Melitaea maturna - / urbanoides / Warn.” // “ZMH 827658”; “Holstein / Innien / [illegible]” // “VON / G. WARNECKE / W. LINZ. HAMB.” // “Sig. W. Linz / Eing. 12 – 74” // “ZMH 827658”.

Original locality

Germany: “Innien bei Neumünster (Holstein).”; Schleswig-Holstein.

Remarks

Warnecke (1942) proposed this name as a “form” of Melitaea maturna (Linnaeus, 1761). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. The title already indicates that this form constitutes an individual aberration and thus an unavailable name.

tenuicula Verity, 1919

Original combination

Melitaea athalia race tenuicula mihi” Verity, 1919 Ent. Rec. 31: 193–194.

Current combination

Melitaea athalia tenuicula Verity, 1919.

Current status

Valid subspecies.

Original material

Labelled as “Type” 6♂♂ (ZMH 833455–833460) (Fig. 110). “tenuicula Vrty” // helvetica Rühl” // “Marche / Bolognola / 28 VI 1937 / Querci.” // [blank label] // “ZMH 833455”; “tenuicula Vrty” // helvetica Rühl” // “Marche / Bolognola / 15 VI 1937 / Querci.” // [blank label] // “ZMH 833456”; “tenuicula Vrty” // “Marche / Bolognola / 17 VI 1937 / Querci.” // [blank label] // “ZMH 833457”; “tenuicula Vrty” // helvetica Rühl” // “Marche / Bolognola / 18 VI 1937 / Querci.” // [blank label] // “ZMH 833458”; “tenuicula Vrty” // “Marche / Bolognola / 28 VI 1937 / Querci.” // [blank label] // “ZMH 833459”; “tenuicula Vrty” // “Marche / Bolognola / 27 VI 1937 / Querci.” // [blank label] // “ZMH 833460”.

Original locality

Italy: “Bolognola in the Sibillini Mountains, 1200 m”.

Remarks

Verity (1919) proposed this name as a “race” of Melitaea athalia (Rottemburg, 1775). According to Kudrna (1983: 83–84), this name is available as subspecific. The year on the specimen labels (1937) indicate specimens were collected 18 years after description (1919) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these six specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens

varissima Verity, 1914

Original combination

Melitaea [parthenie Bkh] varia Meyer-Dür varissima nom. nov.” Verity, 1914 (1913) Boll. Soc. Ent. Ital. 45: 210.

Current combination

Melitaea varia varissima Verity, 1914.

Current status

Valid subspecies.

Original material

Labelled as “Type” 6♂♂ (ZMH 833461–833466) (Fig. 111). “varissima Vrty” // “varia Meyer-Dür” // “Marche / Bolognola / 25.VI.1937 / Querci” // [blank label] // “ZMH 833461”; “varissima Vrty” // “Marche / Bolognola / 20.VI.1937 / Querci” // [blank label] // “ZMH 833462”; “varissima Vrty” // “Marche / Bolognola / 14.VI.1937 / Querci” // [blank label] // “ZMH 833462”; “varissima Vrty” // “Marche / Bolognola / 14.VI.1937 / Querci” // [blank label] // “ZMH 833463”; “varissima Vrty” // “Marche / Bolognola / 5.VI.1937 / Querci” // [blank label] // “ZMH 833464”; “varissima Vrty” // “Marche / Bolognola / 22.VI.1937 / Querci” // [blank label] // “ZMH 833465”; “varissima Vrty” // “Marche / Bolognola / 25.VI.1937 / Querci” // [blank label] // “ZMH 833466”.

Original locality

Italy: Bologna “Nei Faggeti di Bologna, nella località Costarotara e a Piano Astore dei Monti Sibillini, da 1200 a 1600 m. d’altitudine” [Bologna, vicinity of Costarotara and Piano Astore of the Sibillini Mountains, 1200–1600 m.].

Remarks

Verity (1919) proposed this name as a “race” of Melitaea varia Meyer-Dür, 1851. According to Kudrna (1983: 84), this name is available as subspecific. The year on the specimen labels (1937) indicate specimens were collected 23 years after description (1914) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these six specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.

riloensis Züllich, 1936

Original combination

Melitaea cinxia L. var. riloensis Züllich.” Züllich, 1936 Z. Öst. Ent. Ver. 21: 27.

Current combination

Melitaea cinxia riloensis Züllich, 1936.

Current status

Junior subjective synonym of Melitaea cinxia clarissa Staudinger.

Type material

Syntypes 2♂♂ (ZMH 827646–827647) (Fig. 112). “COTYPE / Mel. cinxia L. / ♂ v. riloensis / Z. d. Ö. Ent. V. 1936 / DR. ZÜLLICH, WIEN” // “BULGARIEN JUNI 1923 / RYLA GEB. 1200 bis 1800 m / O. BUBACEK WIEN” // [blank label] // “ZMH 827646”; “COTYPE / Mel. cinxia L. / ♂ v. riloensis / Z. d. Ö. Ent. V. 1936 / DR. ZÜLLICH, WIEN” // “BULGARIEN JUNI 1923 / RYLA GEB. 1200 bis 1800 m / O. BUBACEK WIEN” // [blank label] // “ZMH 827647”.

Type locality

Bulgaria: “auf dem Steilhang östlich des Riloklosters in einer Höhe von etwa 1200 m.” [at a steep slope east of the Rilo monastry at an altitude of about 1200 m]. 1901.

Remarks

Züllich (1936) proposed this name as a “var.” of M. cinxia (Linnaeus, 1758). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.

ab. mediofasciata Bubacek, 1926

Original combination

Melitaea dejone H.G. ab. mediofasciata Bub. ab. nova.” Bubacek, 1926 Verh. Zool.-Bot. Ges. Wien 74–75: (9).

Current combination

Melitaea deione ab. mediofasciata Bubacek, 1926.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 833454) (Fig. 113). “Type / Mel. rondoui / ab. mediofasciata / Otto Bubacek” // “France, Gèdre / Hautes Pyrenées / Juli 1923 / Col. Otto Bubacek” // [blank label] // “ZMH 833454”.

Original locality

France: Pyrenées.

Remarks

Bubacek (1926) proposed this name as an aberration of M. dejone Geyer, 1832. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

ab. johni Stauder, 1922

Original combination

M. dictynna ab. johni” Stauder, 1922 Ent. Anz. 2: 44.

Current combination

Melitaea diamina ab. johni Stauder, 1922.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827648) (Fig. 114). “Lainau 800 m / [illegible] / 1.VII.17 Schawerda” // “Type” // “dictynna f. / johni Stauder” // “Type” // “ZMH 827648”.

Original locality

Austria: “Salzkammergut (Lainautale, Hallstatt)”.

Remarks

Stauder (1922) proposed this name as an aberration of M. dictynna Esper, 1778. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

balcanica Züllich, 1936

Original combination

Melitaea trivia Schiff. var. balcanica Züllich.” Züllich, 1936 Z. Öst. Ent. Ver. 21: 27.

Current combination

Melitaea trivia balcanica Züllich, 1936.

Current status

Junior subjective synonym of Melitaea trivia lathon Fruhstorfer, 1917.

Type material

Syntypes 2♂♂ (ZMH 827663–827664) (Fig. 115). “BULGARIEN JUNI 1928 / RYLA GEB. 1200 bis 1800 m / O. BUBACEK” // [blank label] // “COTYPE / Mel. trivia / ♂ v. balcanica / Z. d. Ö. Ent. V. 1936 / DR. ZÜLLICH, WIEN” // “ZMH 827663”; “BULGARIEN JUNI 1928 / RYLA GEB. 1200 bis 1800 m / O. BUBACEK” // [blank label] // “COTYPE / Mel. trivia / ♂ v. balcanica / Z. d. Ö. Ent. V. 1936 / DR. ZÜLLICH, WIEN” // “ZMH 827664”.

Type locality

Bulgaria: “in den gebirgigen Teilen der zentralen und östlichen Balkanhalbinsel (Krain, Bosnien, Herzegowina etc.)”. [in the mountainous parts of the central and eastern Balkan peninsula].

Remarks

Züllich (1936) proposed this name as a variety of M. trivia ([Denis & Schiffermüller], 1775). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.

catapelioides Stauder, 1918

Original combination

Melitaea trivia catapelioides nov. subspec. m.” Stauder, 1918 Zeit. Wiss. Ins. Biol. 14: 57.

Current combination

Melitaea trivia catapelioides Stauder, 1918.

Current status

Junior subjective synonym of nominotypical Melitaea trivia ([Denis & Schiffermüller], 1775).

Original material

Labelled as “Type” 1♂ (ZMH 827662) (Fig. 116). “Type” // “Luen / Calabria mer. / Aspromonte 950 m. / 6.VII 1920 / H. Stauder legit.” // “trivia catape - / lioides Stauder” // “Type” // “Sig. G. Warnecke / Eing. Nr. 5, 1945” // “ZMH 827662”.

Original locality

Italy: “Unteritalien (Kalabrien)”.

Remarks

The year on the specimen label (1920) indicates specimen was collected two years after description (1918) from the same location (i.e., Italy, Kalabrien) the original holotype material was collected. As a result, the label of the specimen treated as “Type” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, this specimen is erroneously labelled as type, should be annotated as “Not a Type” and is treated here as “Non-type” specimen.

ignasiti de Sagarra, 1926

Original combination

Melitaea trivia Schiff., rassa ignasiti, nova rassa” de Sagarra, 1926 Butl. Inst. Catal. Hist. Nat. (2)6: 131.

Current combination

Melitaea trivia ignasiti de Sagarra, 1926.

Current status

Valid subspecies.

Type material

Paralectotypes 1♂1♀ (ZMH 827660–827661) (Fig. 117). “CATALONIA / Seva 700 m. / 3.8.25 Romei” // “M. triviaignasiti Sag. / co-type ♂ Querci” // “ZMH 827660”; “CATALONIA / Seva 700 m. / 3.8.25 Romei” // “Sammlung / R. Jähnig / Eing. Nr.29,1949” // “M. triviaignasiti Sag. / co-type ♀ Querci” // “L.28” // “ZMH 827661”.

Type locality

Spain: Catalonia, “Martorell”.

Remarks

de Sagarra (1926) proposed this name as a “rassa = race” of M. trivia ([Denis & Schiffermullar], 1775). The term “rassa = race” which was employed by de Sagarra in 1926 was often used in the current sense of subspecies. As a result, the first line of article 45.6 (ICZN 1999) – the rank denoted by a species-group name following a binomen is subspecific – is applied here, it is thus available as subspecific. Higgins (1941) synonymised it with M. trivia pseudodidyma Rebel, 1905 but he reconsidered it as a valid subspecies in his subsequent books. García-Barros et al. (2013) made a comment in Fauna Ibérica that “the orange color of the Iberian specimens is lighter than those from other European areas; those have been attributed to subspecies “ignasiti de Sagarra, 1926”, which, however, does not seem significantly different. Furthermore, it does not seem significantly differentiated from a genetic point of view (Wahlberg pers. comm.).” The taxon ignasiti was reinstated as a valid species by van Oorschot and Coutsis (2014), and this was accepted by Macia et al. (2015) who designated lectotype and paralectotypes. Therefore, specimens at ZMH are by default paralectotypes. However, most other recent authors (e.g., Kudrna 2019) treat this taxon as a synonym to M. trivia, because there is no evidence for species status. Even Oorschot and Coutsis (2014) mention intermediates indicating hybridisation. It should be noted that the species name is not mentioned in the latest European checklist (Wiemers et al. 2018).

Tribe Nymphalini Rafinesque, 1815

guhni Tschauner, 1926

Original combination

Vanessa urticae var. guhni” Tschauner, 1926 Int. Ent. Z. 20: 229, 273.

Current combination

Aglais urticae guhni (Tschauner, 1926).

Current status

Junior subjective synonym of Aglais urticae urticae (Linnaeus, 1758).

Type material

Paratype 1♂ (ZMH 827667) (Fig. 118). “Paratypoid” // “Vanessa urticae / nov. var. guhni / Paratype e. coll. / Tschauner, Berlin 1926” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827667”.

Type locality

Germany: Berlin.

Remarks

This case seems ambiguous, because Tschauner (1926) writes that the new name constitutes a local variation, if not a new race, although he also notes that he is not sure whether the variation is really heritable. On page 273 he names it again as “Vanessa urticae var. Guhni nov. spec.“ [sic!]. See also the critical remark by Kitt (1927).

f. clarior Warnecke, 1959

Original combination

Araschnia levana (Linnaeus) gen. vern. n. f. clarior” Warnecke, 1959 Mitt. Faun. Arb. Gem. Schlesw. Holst. 1 (10.): 53.

Current combination

Araschnia levana f. clarior Warnecke, 1959.

Current status

Infrasubspecific and thus unavailable name.

Original material

Labelled as “Type” 2♂♂ (ZMH 825009–825010) (Fig. 119). “Schleswig / Elsdorfer Sch. / 20.3.97” // “n.f. clarior W. / Coll. Warnecke / Eing. Nr. 5-1949” // “ZMH 825009”; “Schleswig / Elsdorfer Sch. / c. l. 20.3.97” // “n.f. clarior W. / Coll. Warnecke / Eing. Nr. 5-1949” // “ZMH 825010”.

Original locality

Germany: Schleswig-Holstein.

Remarks

Warnecke (1959) proposed this name as an aberration of A. levana (Linnaeus, 1758). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. From the text of the description it is unambiguously clear that the author is describing an aberration and not a subspecies.

ab. marginelineata Horch, 1932

Original combination

Araschnia levana L. gen. aest. prorsa ab. marginelineata” Horch, 1932 Int. Ent. Z. 26:338.

Current combination

Araschnia levana prorsa ab. marginelineata Horch, 1932.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Kotype” 1♂ (ZMH 825011) (Fig. 120). “f. marginelineata / Horch / Gub. I. E Z; 26., 1932/3, / P. 337.” // “Umgeg. v. Hamburg / Radbruch / e. l. 10.7.32 / R. Horch Hamburg” // “Kotype” // “ZMH 825011”.

Original locality

“Lüneburger Heide”, Hamburg [Germany].

Remarks

Horch (1932) proposed this name as an abberation of A. levana (Linnaeus, 1758). This species is known for having two forms, levana and prorsa that represent the spring and summer broods respectively. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

Subfamily Satyrinae Boisduval, 1833

Tribe Satyrini Boisduval, 1833

alphea Warnecke, 1919

Original combination

Satyrus arethusa Schiff. nov. subsp. alphea Warn.” Warnecke, 1919 Int. Ent. Z. 12: 182.

Current combination

Arethusana arethusa alphea (Warnecke, 1919).

Current status

Junior subjective synonym of Arethusana arethusa arethusa ([Denis & Schiffermüller], 1775).

Type material

Syntype 1♂ (ZMH 827676) (Fig. 121). “Type” // nov. v. Alphea” / Warn.” // “Type” // “Griechenld.” // “ZMH 827676”.

Type locality

Greece: “Griechenland, Velucci und Parnass”.

asperomontana Stauder, 1921

Original combination

Satyrus circe L. subsp. n. asperomontana” Stauder, 1921 Dt. Ent. Z. Iris 35: 28.

Current combination

Brintesia circe asperomontana (Stauder, 1921).

Current status

Junior subjective synonym of Brintesia circe circe (Fabricius, 1775).

Type material

Syntype 1♂ (ZMH 827680) (Fig. 122). “1/7 / Calabria mer. / Asperomonte 1200 m. / Polsi 1920 / H. Stauder legit.” // “Type” // “circe / asperomontana / Type” // “ZMH 827680”.

Type locality

Italy: “Polsibeken, Aspromonte 1350–1500 m.”

f. rufobrunnea Warnecke, 1942

Original combination

Coenonympha arcania L. n. f. rufobrunnea.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 102.

Current combination

Coenonympha arcania f. rufobrunnea Warnecke, 1942.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827677) (Fig. 123). “Itzehoe / (Holstein)” // “n.f. rufobrunnea Warn. / Type / Iris 1942, p. 102” // “Katalog Nr. 519 / HL” // “Coenon. arcania - / f. rufobrunnea / Warn. / L.38” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827677”.

Original locality

Germany: Schleswig-Holstein, Itzehoe.

Remarks

Warnecke (1942) proposed this name as a form of C. arcania (Linnaeus, 1761). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. From the text it is clear that the author describes an individual form / aberration.

tyrrhena Stauder, 1915

Original combination

Coenonympha arcania tyrrhena m. subsp. nov.” Stauder, 1915 Zeit. Wiss. Ins. Biol. 11: 1, t. 2, f. 17.

Current combination

Coenonympha arcania tyrrhena Stauder, 1915.

Current status

Junior subjective synonym of Coenonympha arcania arcania (Linnaeus, 1761).

Type material

Syntype 1♂ (ZMH 827761) (Fig. 124). “Type” // “Calabria / Umg. Paola-Co-/Senza 18.6.1920 / H. Stauder” // “Type / v. tyrrhena Stauder” // “arcania v. / tyrrhena Staud.” // “ZMH 827761”.

Type locality

Italy: Monte Martinello bei 1000 m. [Castello Cabiaglio, Province of Varese, Italy].

ab. energica Bubacek, 1923

Original combination

Coenonympha corinna Hb. ab. nova energica Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).

Current combination

Coenonympha corinna ab. energica Bubacek, 1923.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827740) (Fig. 125). “Type / Coen. corinna Hb. / ab. energica Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827740”.

Original locality

France: Korsika [Corse].

Remarks

Bubacek (1923) proposed this name as an aberration of C. corinna (Hübner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

ab. caeca Bubacek, 1923

Original combination

Coenonympha corinna Hb. ab. nova caeca Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).

Current combination

Coenonympha corinna ab. caeca Bubacek, 1923.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827741) (Fig. 126). “Type / Coen. corinna Hb. / ab. caeca Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827741”.

Original locality

France: Korsika [Corse].

Remarks

Bubacek (1923) proposed this name as an aberration of C. corinna (Hübner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

ab. anophthalmica Bubacek, 1923

Original combination

Coenonympha corinna Hb. ab. nova anophthalmica Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).

Current combination

Coenonympha corinna ab. anophthalmica Bubacek, 1923.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827742) (Fig. 127). “Type / Coen. corinna Hb. / ab. anophthalmica Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827742”.

Original locality

France: Korsika [Corse].

Remarks

Bubacek (1923) proposed this name as an aberration of C. corinna (Hübner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

ab. saturata Bubacek, 1923

Original combination

Coenonympha corinna Hb. ab. nova saturata Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).

Current combination

Coenonympha corinna ab. saturata Bubacek, 1923.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827743) (Fig. 128). “Type / Coen. corinna Hb. / ab. saturata Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827743”.

Original locality

France: Korsika [Corse].

Remarks

Bubacek (1923) proposed this name as an aberration of C. corinna (Hübner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

ab. nigricans Bubacek, 1923

Original combination

Coenonympha corinna Hb. ab. nova nigricans Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).

Current combination

Coenonympha corinna ab. nigricans Bubacek, 1923.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827744) (Fig. 129). “Type / Coen. corinna Hb. / ab. nigricans Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827744”.

Original locality

France: Korsika [Corse].

Remarks

Bubacek (1923) proposed this name as an aberration of C. corinna (Hübner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

ab. macrophthalmica Bubacek, 1923

Original combination

Coenonympha corinna Hb. ab. nova macrophthalmica Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).

Current combination

Coenonympha corinna ab. macrophthalmica Bubacek, 1923.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 2♂♂ (ZMH 827745–827746) (Fig. 130). “Type / Coen. corinna Hb. / ab. macrophthalmica Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827745”; “Type / Coen. corinna Hb. / ab. macrophthalmica Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827746”.

Original locality

France: Korsika [Corse].

Remarks

Bubacek (1923) proposed this name as an aberration of C. corinna (Hübner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

ab. gynandra Bubacek, 1923

Original combination

Coenonympha corinna Hb. ab. nova gynandra Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).

Current combination

Coenonympha corinna ab. gynandra Bubacek, 1923.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 2♂♂ (ZMH 827747–827748) (Fig. 131). “Type / Coen. corinna Hb. / ab. gynandra Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827747”; “Type / Coen. corinna Hb. / ab. gynandra Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827748”.

Original locality

France: Korsika [Corse].

Remarks

Bubacek (1923) proposed this name as an aberration of C. corinna (Hübner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

f. pulcherrima Warnecke, 1942

Original combination

Coenonympha iphis Schiff., ♀ n.f. pulcherrima.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 102.

Current combination

Coenonympha glycerion f. pulcherrima Warnecke, 1942.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♀ (ZMH 827738) (Fig. 132). “Mölln i. lotz / 30.6.35 [illegible]” // “30.6.35 / Molln” // “♀ n.f. pulcherrima / Warn. Type / Iris 1942, 102” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “Type” // “ZMH 827738”.

Original locality

Germany: “Mölln (Lauenburg)“; Schleswig-Holstein, Herzogtum Lauenburg, Mölln.

Remarks

Warnecke (1942) proposed this name as a form of C. iphis ([Denis & Schiffermüller], 1775). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. From the text it is clear that the author describes an aberration and not a subspecies.

ab. oikeia Dannehl, 1927

Original combination

Coenonympha iphis Schiff. ab. oikeia Dhl.” Dannehl, 1927 Mitt. Münch. Ent. Ges. 17: 5.

Current combination

Coenonympha glycerion ab. oikeia Dannehl, 1927.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 835591) (Fig. 133). “25.6” // “Italia central. / Mti. Simbruini / coll. F. Dannehl” // “oikeia / Dhl. / Cotype. / G.” // “Cotype” // “25.6. 1926 / Mti. Simbruini / Mittel.Italien” // “ZMH 835591”.

Original locality

Italy: “Mt. Simbruini von etwa 1400 m Höhe”.

Remarks

Dannehl (1927) proposed this name as an aberration of C. iphis ([Denis & Schiffermüller], 1775). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

iranica Schwingenschuss, 1939

Original combination

Coenonympha leander Esp. nov. ssp. iranica Sch.” Schwingenschuss, 1939 Ent. Z. 53: 62.

Current combination

Coenonympha leander iranica Schwingenschuss, 1939.

Current status

Valid subspecies.

Type material

Syntype 1♂ (ZMH 827714) (Fig. 134). “Pers. Elbursgeb. / Kendevanpass / 2800 m, 3.-8.7.36 / Schwingenschuss” // “Cotypus” // “ssp. iranica Schwing.” // “Coll. Bytinski-Salz / Eing. Nr. 20 – 60” // “ZMH 827714”.

Type locality

Iran: “Am Kendevan (Sch., W., Pf.) von 2600 m bis 3000 m” [Kandovan pass].

rhenana Gradl, 1933

Original combination

Coenonympha oedipus F. subspecies rhenana” Gradl, 1933 Int. Ent. Z. 27: 261.

Current combination

Coenonympha oedippus rhenana Gradl, 1933.

Current status

Junior subjective synonym of nominotypical Coenonympha oedippus (Fabricius, 1787).

Type material

Syntypes 4♂♂ (ZMH 827722–827725) (Fig. 135). “Feldk / Vorarlberg / Prof. F. Gradl / 31.7.32 / Rheinau” // “Subsp. / rhenana / F-Gradl” // “Cotype” // “L.23 / Coenonympha / oedippus - / rhenana Gradl” // “Sig. G. Warnecke / Eing Nr 5, 1949” // “ZMH 827722”; “Vorarlberg / Prof. F. Gradl / Rheinauen / 12.7.27” // “Subsp. / rhenana / F-Gradl” // “Cotype” // “ZMH 827723”; “Vorarlberg / Prof. F. Gradl / Feldkirch. Rhein / auen. 12.7.31” // “Subsp. / rhenana / F-Gradl” // “Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827724”; “Vorarlberg / Prof. F. Gradl / Feldk-Rheinau / 28.7.32” // “Subsp. / rhenana / F-Gradl” // “Co-Type” // “Type” // “ZMH 827725”.

Type locality

Germany: “in den Niederungen des Rheintales”. [in the lower region of the Rhine valley].

galvagnii Stauder, 1923

Original combination

C. p. galvagnii m. nova subspec. (partim aberr.)” Stauder, 1923 Zeit. Wiss. Ins. Biol. 18: 59, t. 3, f. 6–13.

Current combination

Coenonympha pamphilus galvagnii Stauder, 1923.

Current status

Junior subjective synonym of nominotypical Coenonympha pamphilus (Linnaeus, 1758).

Type material

Syntype 1♂ (ZMH 827739) (Fig. 136). “Type” / “H. Stauder / Litorale austr. / Triest Umgbg. / 12/8/1913” // “pamphilus / galvagnii Staud.” // “Type” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827739”.

Type locality

Italy: Bologna “Bei Grado auf den Lagunen in der g. v. und sonst im Küstengebiete, bei Triest, Görz.” [Bologna, vicinity of Costarotara and Piano Astore of the Sibillini Mountains, 1200–1600].

f. maculata Warnecke, 1942

Original combination

Coenonympha tiphon Rott., ♀, n. f. maculata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.

Current combination

Coenonympha tullia f. maculata Warnecke, 1942.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 2♀♀ (ZMH 827762–827763) (Fig. 137). “Bahrenfeld / 1913” // “♀ n.f. maculata / Warn. Cotype / Iris 1942, 103” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827762”; “Segeberg / Juli 13” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827763”.

Original locality

Unclear; Germany: Bahrenfeld, Hamburg “aus Holstein.”; publication states: Holstein, label of cotype specimen from Warnecke states Bahrenfeld, ?, (one further specimen without type label: Segeberg)

Remarks

Warnecke (1942) proposed this name as a form of C. tiphon (Rottemburg, 1775). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. From the text and already from the title of the paper it is clear that the author describes an aberration. Papilio tiphon Rottemburg, 1775 is junior subjective synonym of Coenonympha tullia (Müller, 1764).

flavofasciata Heyne, 1895

Original combination

Erebia flavofasciata Heyne.” Heyne, [1895]; in Rühl & Heyne, Die Pal. Gross-Schmett. 1: 805.

Current combination

Erebia flavofasciata Heyne, 1895.

Current status

Valid species.

Type material

Paratype 1♂ (ZMH 833469) (Fig. 138). “Campolungo / Pass / Oberst v. Nolte” // “Flavofasciata” // “Type oder Paratype / e coll. von Nolte / s. Heyne-Rühl, 1895, 805, / der keine Samml. hatte. / von Fuchs-Hbg ded.” // “ZMH 833469”.

Type locality

Italy: Tessin, Campolungo pass close to Fusio.

peneplana Berger, 1936

Original combination

Erebia aethiops ESPER, race peneplana, nov. race” Berger, 1936 Lambillionea 36: 199.

Current combination

Erebia aethiops peneplana Berger, 1936.

Current status

Valid subspecies.

Type material

Paratypes 3♂♂ (ZMH 835588–835590) (Fig. 139). “PARATYPE / Collection / Luc. Berger” // “Tellin. (R.d. Boyes) / BELGIQUE / 5/8/1936 / Coll. Luc. BERGER.” // [blank label] // “ZMH835588”; “PARATYPE / Collection / Luc. Berger” // “Tellin. (R.d. Boyes) / BELGIQUE / 5/8/1936 / Coll. Luc. BERGER.” // [blank label] // “ZMH835589”; “PARATYPE / Collection / Luc. Berger” // “Tellin. (R.d. Boyes) / BELGIQUE / 16/8/1934 / Coll. Luc. BERGER.” // [blank label] // “ZMH835590”.

Type locality

Belgium: “env. de Han-sur-Lesse”.

clorinda Hartig, 1940

Original combination

Erebia epiphron Fn. ssp. clorinda n.” Hartig, 1940 Mem. Soc. Ent. Ital. 18: 186.

Current combination

Erebia epiphron clorinda Hartig, 1940.

Current status

Junior subjective synonym of Erebia epiphron amplevittata Verity, 1921.

Type material

Paratypes 1♂1♀ (ZMH 827718–827719) (Fig. 140). “Paratypus ♀ / Er. epiphr. / clorinda / Hartig” // “1700 m / Marche / Bolognola / 20.VI 1937 / Querci” // [blank label] // “ZMH 827718”; “Paratypus ♀ / Er. epiphr. / clorinda / Hartig” // “1700 m / Marche / Bolognola / 20.VI 1937 / Querci” // [blank label] // “ZMH 827718”; “Paratypus ♂ / Er. epiphr. / clorinda Hartig” // “1700 m / Marche / Bolognola / 20.VI 1937 / Querci” // “[blank label] // “ZMH 827719”.

Type locality

Italy: “Bolognola”.

vogesiaca Goltz, 1914

Original combination

Erebia epiphron vogesiaca” Goltz, 1914 Dt. Ent. Z. Iris 28:107–109.

Current combination

Erebia epiphron vogesiaca Goltz, 1914.

Current status

Junior subjective synonym of Erebia epiphron mackeri Fuchs, 1914).

Type material

Syntype 1♂ (ZMH 827679) (Fig. 141). “COTYPE / vogesiaca / v.d.Goltz” // “VII 1913 / Fauxkopf / Vogesen” // “Fauxkopf / Juli 1913 / Kesenheimer” // “ZMH 827679”.

Type locality

France: “Vogesen”.

leonhardi Fruhstorfer, 1918

Original combination

Erebia melas leonhardi subspec. nova” Fruhstorfer, 1918 Arch. Naturgesch. 84 A7: 131.

Current combination

Erebia melas leonhardi Fruhstorfer, 1918.

Current status

Valid subspecies.

Type material

Syntypes 11♂♀ (ZMH 827727–827737) (Fig. 142). “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827727”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827728”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827729”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827730”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827731”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827732”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827733”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827734”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827735”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827736”; “Velebit, Ostaria” // “Cotypus” // “melas leonhardi Cotype / Velebit Ostria 19[..]” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827737”.

Type locality

Croatia: “Velebit, Ostaria”.

paradisi Dannehl, 1929

Original combination

Erebia stygne O. paradisi Dhl.” Dannehl, 1929 Mitt. Münch. Ent. Ges. 19: 98.

Current combination

Erebia meolans paradisi Dannehl, 1929.

Current status

Junior subjective synonym of nominotypical Erebia meolans (de Prunner, 1798).

Type material

Syntypes 3♂♂ (ZMH 835585–835587) (Fig. 143). “19.6” / “Apenn. merid. / Mt. Paradiso / 1500–2000 m / coll. F. Dannehl” // “19.6. 1928 / Mte. Paradiso / Central – Apenninen” // “ZMH 835585”; “22.6” / “Apenn. merid. / Mt. Paradiso / 1500–2000 m / coll. F. Dannehl” // “20.6. 1928 / Mte. Paradiso / Central – Apenninen” // “ZMH 835586”; “19.6” / “Apenn. merid. / Mt. Paradiso / 1500–2000 m / coll. F. Dannehl” // “19.6. 1928 / Mte. Paradiso / Central – Apenninen” // “ZMH 835587”.

Type locality

Italy: “Monte Paradiso 1600–1900 m, Monte Rotella, Montagna Grande 1800–2100 m (südl. Abruzzen).”

benacensis Warren, 1933

Original combination

Erebia ottomana ssp. benacensis nov.” Warren, 1933 Ent. Rec. 45: 40.

Current combination

Erebia ottomana benacensis Warren, 1933.

Current status

Junior subjective synonym of nominotypical Erebia ottomana Herrich-Schäffer, (1847).

Type material

Syntypes 5♂♂ (ZMH 833449–833453) (Fig. 144). “4.7” // “Teriolis merid. / Monte Baldo / 2000 m. / coll. F. Dannehl” // “4.7. 1930 / Mte. Baldo 2000 m / (Gardasee)” // “ZMH 833449”; “4.7” // “Teriolis merid. / Monte Baldo / 2000 m. / coll. F. Dannehl” // “4.7. 1930 / Mte. Baldo 2000 m / (Gardasee)” // “ZMH 833450”; “5.7” // “Teriolis merid. / Monte Baldo / 2000 m. / coll. F. Dannehl” // “5.7. 1930 / Mte. Baldo 2000 m / (Gardasee)” // “ZMH 833451”.

Type locality

Italy: “Mte. Baldo near Lake Garda”.

kotzschae Goltz, 1937

Original combination

Erebia kotzschae” Goltz, 1937 Ent. Rdsch. 54: 363.

Current combination

Paralasa kotzschae (Goltz, 1937).

Current status

Valid species.

Type material

Syntype 1♂ (ZMH 827715) (Fig. 145). “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500–4000 m Mitte Juli / leg. H.&E. Kotzsch” // “COTYPE / kotzschae / v.d. Goltz” // “Kotschi v.d. Goltz” // “Coll. Bytinski-Salz / Eing Nr 20 – 60” // “ZMH 827715”.

Type locality

Afghanistan: Hindukush.

maxima Bang-Haas, 1933

Original combination

Satyrus autonoe maxima O. B.-Haas, subsp. nov.” Bang-Haas, 1933 Ent. Z. 47: 98.

Current combination

Hipparchia autonoe maxima (Bang-Haas, 1933).

Current status

Valid subspecies.

Type material

Syntypes 2♂♂ (ZMH 827681–827682) (Fig. 146). “Kansu mer. / Min-chan / Tanho / 2500 m. Juli” // “Co-Type / O. Bang-Haas” // “L.13 / Sat. autonoë - / maxima / O.B.H.” // “Sammlung / R. Jähnig / Eing Nr. 29, 1949” // “ZMH 827681”; “Kansu mer. or. / Fukiang / Peilingschan sept. / 2500 m. Juli” // “Co-Type / O. Bang-Haas” // “L.14 / Sat. autonoë - / maxima / O.B.H.” // “Sammlung / R. Jähnig / Eing Nr. 29, 1949” // “ZMH 827682”.

Type locality

China: “Kansu mer., Lihsien, Kialing Fluß, 2500 m.”

persiscana Verity, 1937

Original combination

Eumenis allionii, G.-H. = fatua Freyer, persiscana, nom. nov.” Verity, 1937 Ent. Rec. 49: 100.

Current combination

Hipparchia fatua persiscana (Verity, 1937).

Current status

Valid subspecies.

Type material

Syntypes 31♂♂ (ZMH 827683–827713) (Fig. 147). “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827683”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827684”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827685”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827686”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827687”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827688”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827689”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827690”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827691”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827692”; “Iran / Keredj 1800 m / 21.8.1936 / Brandt” // “Cotypus” // “ssp. persicana / Vrty” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827693”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827694”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827695”; “Iran / Keredj 1800 m / 25/8 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827696”; “Iran / Keredj 1800 m / 25/8 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827697”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827698”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827699”; “Iran / Keredj 1800 m / 25/8 1936 / Brandt” // “201” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827700”; “Iran / Keredj 1800 m / 21/8 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827701”; “Iran / Keredj 1200 m / 1936 / Brandt” // “1600 25/VIII” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827702”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827703”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827704”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827705”; “Iran / Keredj (1200 m.) / 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827706”; “Iran / Keredj 1600 m / 25/8 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827707”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827708”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827709”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827710”; “Iran / Keredj 1200 m / 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827711”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827712”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827713”.

Type locality

Iran: Tehran, “Keredj [Karaj], in the Elburz Mountains, at m. 1500 to 1600”.

ab. pallida Bubacek, 1923

Original combination

Satyrus neomiris God. ab. nova pallida Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).

Current combination

Hipparchia neomiris ab. pallida (Bubacek, 1923).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827726) (Fig. 148). “Type / Sat. neomiris God / ab. pallida Bub. / Otto Bubacek Z.BG. 1922” // “Col de Verg. Cor- / se Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827726”.

Original locality

France: Corse, Col de Verg.

Remarks

Bubacek (1923) proposed this name as an aberration of S. neomiris (Godart, 1824). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

albinervata Warnecke, 1944

Original combination

“[Satyrus semele L.] n. ab. albinervata” Warnecke, 1944 Dt. Ent. Z. Iris 57: 40.

Current combination

Hipparchia semele ab. albinervata (Warnecke, 1944).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Holotype” 1♀ (ZMH 827749) (Fig. 149). “Insel Sylt / Kampen / VIII 1942” // “n.f. albinervata / Warnecke / Type” // “Holotype” // “Sammlung G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827749”.

Original locality

Germany: “Sylt, Kampen”.

Remarks

Warnecke (1944) proposed this name as a form of S. semele (Linnaeus, 1758). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

siviae Fuchs, 1954

Original combination

Lymanopoda shefteli siviae” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84.

Current combination

Lymanopoda shefteli siviae Fuchs, 1954.

Current status

Valid subspecies.

Type material

Lectotype 1♂ (ZMH 827764) and 3 ♂♂ paralectotypes (ZMH 827765–827767) (Fig. 150). “Lymnopoda / siviae / v. Fuchs / Cordillere von / Sivia, Ayacucho / 2500 m.” // “Type” // “Holotype / Lymnopoda siviae / von Fuchs” // “Lymnopoda galactea / shefteli Dyar / G. Lamas. det. 1978” // “Lectotype ♂ / Lymnopoda / siviae Fuchs / by G. Lamas, 1978” // “Sammlung / W. v. Fuchs / Eing Nr. 4, 1956” // “ZMH 827764”; “Rio Piene / ca. 2600 m / 7/1920 / Type” // “285 / galactea ab. / Rio Piene 2600 m. 7/20” // “ZMH 82765”; “Rio Piene / ca. 2600 m / 7/1920 / Type” // “ZMH 82766”; “Rio Piene / ca. 2600 m / 7/1920 / Type” // “ZMH 82767”.

Type locality

Peru: “… oberhalb Sivia gefangen, [above Sivia] 2500–3000 m”.

cocuzzana Stauder, 1914

Original combination

Melanargia arge Sulz. (=amphitrite Hb.) cocuzzana m., subspec. nova” Stauder, 1914 Zeit. Wiss. Ins. Biol. 10: 375, f. 12–15.

Current combination

Melanargia arge cocuzzana Stauder, 1914.

Current status

Junior subjective synonym of nominotypical Melanargia arge (Sulzer, 1776).

Type material

Paratypes 3♂♂ (ZMH 827673–827675) (Fig. 151). “Calabria / Umg. Paola-Co-senza 4.6.1913 / H. Stauder” // “arge cocuzzana Stau- /der Paratype” // “ZMH 827673”; “Calabria / Umg. Paola-Co-senza 4.6.1913 / H. Stauder” // “cocuzzana” // “arge cocuzza- / na Stauder / Paratype” // “ZMH 827674”; “Ital. mer. / pen.-Surrentina / 8.6.1913 / H. Stauder” // “Arge cocuzzana / Stauder” // “arge cocuzzana Stau- / der Paratype” // “ZMH 827675”.

Type locality

Italy: vicinity of Paola, Monte Cocuzzo “ an den Hängen [slopes of] des Monte Martinello und Cocuzzo bei Paola”.

ab. huenei Hirschke, 1910

Original combination

Pararge hiera var. ominata Krul. ab. Huenei” Hirschke, 1910 Verh. Zool. Bot. Ges. Wien 60: 411.

Current combination

Lasiommata petropolitana ominata ab. huenei (Hirschke, 1910).

Current status

Infrasubspecific and hence unavailable name..

Original material

Labelled as “Type” 1♂ (ZMH 827717) (Fig. 152). “Estonia / Lechts / FR. V. HUENE” // “Type / Verh. z.b.g. / LX. 1910” // “Pararge hiera / var. ominata / ab. huenei / Hirschke / Estland” // “Coll. Bytinski-Salz / Eing. Nr. 20 – 60” // “ZMH 827717”.

Original locality

Estonia: “aus Esthland”.

Remarks

Hirschke (1910) proposed this name as an aberration of P. h. ominata Krulikowsky, 1903, therefore, as stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable.

felix Warnecke, 1929

Original combination

Pararge felix n. spec. Warn.” Warnecke, 1929 Int. Ent. Z. 22: 365.

Current combination

Lasiommata felix (Warnecke, 1929).

Current status

Valid species.

Type material

Paratype 1♀ (ZMH 827721) (Fig. 153). “S.W. Arabia / Yemen, Sanáa / II.1928. Expedi- / tion Rathjens” // “28.2. /Sanaa” // “Pararge felix / Warnecke 1929 / ♀ Paratype” // “Para-Type” // “Pararge maera - / felix Warn.” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827721”.

Type locality

“Yemen, Sanaa”.

ab. diluta Bubacek, 1923

Original combination

Pararge aegeria L. ab. nova diluta Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).

Current combination

Pararge aegeria ab. diluta Bubacek, 1923.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827678) (Fig. 154) “Type / Par. aegeria L. / ab. diluta Bub. / Otto Bubacek ZBG 1922” // [blank label] // “Evisa, Corse / Juli 1921 / Col. O. Bubacek” // “ZMH 827678”.

Original locality

France: Korsika [Corse].

Remarks

Bubacek (1923) proposed this name as an aberration of P. aegeria (Linnaeus, 1758), therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.

f. albiplaga Warnecke, 1942

Original combination

Pararge algeria L. subsp. egerides Stdgr. n. f. albiplaga ♀.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 102.

Current combination

Pararge aegeria egerides f. albiplaga Warnecke, 1942.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♀ (ZMH 827668) (Fig. 155). “Holstein / Segeberg / 2.-10.8.35” // “n.f. albiplaga / Warn. Type / Iris 1942, 102” // “Type” // “ZMH 827668”.

Original locality

Germany: Holstein [label indicates Segeberg].

Remarks

Warnecke (1942) proposed this name as a form of P. a. egerides (Staudinger, 1871), therefore, as stated by article 45.6.1 it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. In the original description the name “aegeria” is wrongly spelled “algeria” (M. Wiemers pers. comm.).

ab. hermini Hirschke, 1910

Original combination

Pararge megera var. lyssa B. ab. Hermini” Hirschke, 1910 Verh. Zool. Bot. Ges. Wien 60: 411.

Current combination

Lasiommata megera lyssa ab. hermini (Hirschke, 1910).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 827720) (Fig. 156). “Hirschke.Wien / Zara / Mai” // “Type / Verh. Z.G.B. / LX, 1910” // “Pararge megera / var. Lyssa ab. / hermini Hirschke / Dalmatien 5.1909” // “Coll. Bytinski-Salz / Eing Nr 20 – 60” // “ZMH 827720”.

Original locality

Croatia: Dalmatia, vicinity of Gravosa and Zara.

Remarks

Hirschke (1910) proposed this name as an aberration of P. m. lyssa “B.” (sic, recte Hübner, 1829), therefore, as stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable.

picota Fuchs, 1954

Original combination

Argyrophorus picota” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84.

Current combination

Punargentus angusta picota (Fuchs, 1954).

Current status

Valid subspecies.

Type material

Lectotype 1♂ (ZMH 827750) (Fig. 157) 7♂1♀ paralectotypes (ZMH 827751–827758). “Argyrophorus / picota ♂ / v. Fuchs / Cordillere von / Ayacucho / 4500 m” // “Punargentus lamna / angusta Weymer / det. W. Heinrich ♂” // “Type” // “Lectotype ♂ / Argyrophorus / picota Fuchs / by G. Lamas, 1978” // “Sammlung / W. v. Fuchs / Eing. Nr. 4, 1956” // “ZMH 827750”; “Punargentus lamna / angusta Weymer / det. W. Heinrich ♂” // Argyrophorus / picota ♂ / v. Fuchs / Cordillere von / Ayacucho / 4500 m” // “Type” // “Sammlung / W. v. Fuchs / Eing. Nr. 4, 1956” // “ZMH 827751”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta Weymer / det. W. Heinrich ♂” // Coll. H. Rödinger / Eing. Nr. 30, 52” // ZMH 827752”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta Weymer / det. W. Heinrich ♂” // Coll. H. Rödinger / Eing. Nr. 30, 52” // ZMH 827753”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta Weymer / det. W. Heinrich ♂” // “133 / angusta 8/20 / Ayacucho 4/5000 m” // “Coll. H. Rödinger / Eing. Nr. 30, 52” // ZMH 827754”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta Weymer / det. W. Heinrich ♂” // Coll. H. Rödinger / Eing. Nr. 30, 52” // ZMH 827756”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta Weymer / det. W. Heinrich ♂” // Coll. H. Rödinger / Eing. Nr. 30, 52” // ZMH 827757”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta Weymer / det. W. Heinrich ♀” // Coll. H. Rödinger / Eing. Nr. 30, 52” // ZMH 827758”.

Type locality

Peru: near Ayacucho “… Kodillere bei Ayacucho, nur in Höhen über 4000 m, bis 5000 m” [Cordillere close to Ayacucho, only at altitudes above 4000 m, to 5000 m].

arminii Stauder, 1916

Original combination

Epinephele ida arminii subsp. nova.” Stauder, 1916 Zeit. Wiss. Ins. Biol. 12: 62.

Current combination

Pyronia cecilia arminii (Stauder, 1916).

Current status

Junior subjective synonym of Pyronia cecilia cecilia (Vallantin, 1894).

Type material

Syntype 1♂ (ZMH 827716) (Fig. 158). “Type” // “Calabria mer. / Aspromonte 1600 m. / 26.6.1920 / H. Stauder legit.” // “Ida Arminii / Stauder” // “Type” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH Nr. 5, 1949”.

Type locality

Italy: Calabria.

liupiuschani Bang-Haas, 1933

Original combination

Satyrus actaea liupiuschani O. B.-Haas, subsp. nov.” Bang-Haas, 1933 Ent. Z. 47: 99.

Current combination

Satyrus ferula liupiuschani Bang-Haas, 1933.

Current status

Junior subjective synonym of Satyrus ferula ganssuensis Grum-Grshimailo, 1893.

Type material

Syntype 1♂ (ZMH 827669) (Fig. 159). “Kansu mer. / Tsing-schui / Liu-pin-schan / 2000 m. Juli” // “Co-Type / O. Bang-Haas” // “L.17 / Sat. actaea - / liupiuschani O.B.H.” // “Sammlung / R. Jähnig / Eing. Nr.29,1949” // “ZMH 827669”.

Type locality

China: “Kansu mer., Tsingschui, Liu Piu Schan, 2000 m”.

kintschouensis Bang-Haas, 1939

Original combination

Triphysa phryne kintschouensis O. Bang-Haas, subsp. nov.” Bang-Haas, 1939 Dt. Ent. Z. Iris 53: 53.

Current combination

Coenonympha phryne kintschouensis (Bang-Haas, 1939).

Current status

Junior subjective synonym of Coenonympha dohrnii nervosa Motschulsky, 1866.

Type material

Syntypes 2♂♂ (ZMH 827759–827760) (Fig. 160). “18.” // “Mandschuria mer. occ. / Prov. Fengtien / Kintschou / 100 m. April” // “Co-Type / O. Bang-Haas” // “Phryne / v. kintschouensis / O.B.H.” // “L.21 / Triphysa phyrne - / kintschouensis / O.B.-Haas” // “Sammlung R. Jähnig / Eing Nr. 29, 1949” // “ZMH 827759”; “17.” // “Mandschuria mer. occ. / Prov. Fengtien / Kintschou // “Co-Type / O. Bang-Haas” // “L.22 / Triphysa phyrne - / kintschouensis / O.B.-Haas” // “Sammlung R. Jähnig / Eing Nr. 29, 1949” // “ZMH 827760”.

Type locality

China: “Manchuria merid., Prov. Fengtien, Kintschou, 100 m.”.

Unpublished, unavailable names

minor Meinhard

Original combination

Melanargia galathea v. sakaria ab. minor Meinhard.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 835597) (Fig. 161). “Type” // 7/7 1920 / Göttingen / Warteberg” // “ZMH 835597”.

Remarks

This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable.

ab. caeca Dannehl

Original combination

Epinephele ida ab. caeca Dannehl.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Cotype” 1♂ (ZMH 833467) (Fig. 162). “1.6” // Sicilia / Mt. Cuccio / coll. Dannehl” // “Cotype / caeca / Dhl” // “Cotype.” // “1.6 1932 / Mt. Cuccio, / -Sizilien-” // “ZMH833467”.

Remarks

This taxon cannot be found in the literature. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable. The current name of the species is Pyronia cecilia (Vallantin, 1894).

achaltekkensis Holik

Original combination

Satyrus actaea achaltekkensis Holik.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Cotypes” 2♂♂ (ZMH 827670–827671) (Fig. 163). “Transcaspia / Jablonowka / Achal Tekke / 2000 m. Juli” // “Co-Type” // “L.18 / Sat. actaea - / achaltekkensis Hol.” // “Sammlung / R. Jähnig / Eing. Nr. 29, 1949” // “ZMH 827670”; “Transcaspia / Jablonowka / Achal Tekke / 2000 m. Juli” // “Co-Type” // “L.19 / Sat. actaea - / achaltekkensis Hol.” // “Sammlung / R. Jähnig / Eing. Nr. 29, 1949” // “ZMH 827671”.

Original locality

Turkmenistan, Achal Tekke.

Remarks

This taxon cannot be found in the literature. The current name is Satyrus iranica daubi Gross & Ebert, 1975.

rossarmenia Bang-Haas

Original combination

Satyrus actaea rossarmenia Bang-Haas.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Cotype” 1♂ (ZMH 827672) (Fig. 164). “Kagysman / Russ. Armen” // “Co-Type” // “L.16 / Sat. actaea / rossarmenia ?” // “Sammlung R. Jähnig / Eing. Nr.29, 1949” // “ZMH 827672”.

Remarks

This taxon cannot be found in the literature. The current name is Satyrus amasina Staudinger, 1861.

Family Riodinidae Grote, 1895

Subfamily Nemeobiinae Bates 1868

Unpublished, unavailable names

ab. friesi Meinhard

Original combination

Hamearis lucina ab. friesi Meinhard.

Current status

“in litteris” name and hence not available.

Original material

Labelled as “Type” 1♂ (ZMH 835594) (Fig. 165). “Type” // “ab. friesi Mein.” // “10/5 1911 / Göttingen / Hainberg” // “ZMH 835594”.

Remarks

This taxon cannot be found in the literature. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable. The current name is Hamearis lucina (Linnaeus, 1758).

Family Hesperiidae Latreille, 1809

Subfamily Pyrginae Burmeister, 1878

Tribe Carcharodini Verity, 1940

wissmanni Warnecke, 1934

Original combination

Carcharodus swinhoei Wats. nov. subsp. Wissmanni Warn.” Warnecke, 1934 Mitt. Münch. Ent. Ges. 24: 20.

Current combination

Carcharodus alceae wissmanni Warnecke, 1934.

Current status

Valid subspecies.

Type material

Paratypes 5♂♂ (ZMH 827637–ZMH 827638, ZMH 827640–ZMH 827642) (Fig. 166). “S. W. Arabien / Sana’â (Yemen) / 27.2.28 v. Wissm.” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “Carcharodus / Swinhoei Wats. / subsp. wissmanni / Warn. / Cotype” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827637”; “Alceae Swinhoei Wats. / S.W. Arabia / Sana’â (Yemen) 27.2.28 / Rathjens - v. Wissmann” // “Carcharodus / Swinhoei Wats. / subsp. wissmanni / Warn. / Cotype” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827638”; “Süd-Arabien / Yemen: San’aá / 8.8.31 Rathjens” // “Paratype” // Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827640”; “Südwestarabien / Yemen: San’aá / Rathjens 24.6.31” // “Paratype” // Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827641”; “Südwestarabien / Yemen: San’aá / Rathjens 24.7.31” // “Paratype” // “ZMH 827642”.

Type locality

Yemen: Sanaa “Südwest-Arabien, Sana’â (Yemen)”

v. postesseloides Verity, 1938

Original combination

Spialia orbifer, Hüb., race tesselloides, H.-S., II gen. postesseloides, nom. nov.” Verity, 1938 Ent. Rec. 50 Suppl. (4).

Current combination

Spialia orbifer orbifer v. postesseloides Verity, 1938.

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 4♂♂ (ZMH 833498–833501) (Fig. 167). “postesseloides Vrty” // “Macedonia / Olympus, 2500’ / Aug 24, 1935 / Romei” // [blank label] // “ZMH 833498”; “postesseloides Vrty” // “Macedonia / Olympus, 2500’ / July 25, 1936 / Romei” // [blank label] // “ZMH 833499”; “postesseloides Vrty” // “Macedonia / Olympus, 2500’ / July 6, 1936 / Romei” // [blank label] // “ZMH 833500”; “postesseloides Vrty” // “Macedonia / Olympus, 2000’ / July 8, 1936 / Romei” // [blank label] // “ZMH 833501”.

Original locality

Greece: vicinity of Skala.

Remarks

Verity (1938) proposed this name as “nom. nov.“ of S. orbifer tesselloides (Herrich-Schäffer, 1845), therefore, as stated by article 45.6.1 it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. Spialia orbifer tesselloides is currently regarded as a junior subjective synonym of the nominate subspecies.

Tribe Pyrgini Burmeister, 1878

centralitaliae Verity, 1920

Original combination

Hesperia alveus, Hüb., race centralitaliae, mihi” Verity, 1920 Ent. Rec. 32: 4.

Current combination

Pyrgus alveus centralitaliae (Verity, 1920).

Current status

Valid subspecies.

Original material

Labelled as “Type” 4♂♂ (ZMH 833502–833505) (Fig. 168). “centralitaliae Vrty” // “Marche / Bolognola / 6.VII.1937 / Querci” // [blank label] // “ZMH 833502”; “centralitaliae Vrty” // “Marche / Bolognola / 13.VII.1937 / Querci” // [blank label] // “ZMH 833503”; “centralitaliae Vrty” // “Marche / Bolognola / 22.VI.1937 / Querci” // [blank label] // “ZMH 833504”; “centralitaliae Vrty” // “Marche / Bolognola / 27.VI.1937 / Querci” // [blank label] // “ZMH 833505”.

Original locality

Italy: Bologna.

Remarks

Verity (1920) proposed this name as a “race” of H. alveus Hübner, [1803]. As it was given in trinomy and the content of the description clearly indicates that a geographical race was meant, this name is available as subspecific (article 45.6 ICZN 1999). The year on the specimen labels (1937) indicate specimens were collected 17 years after description (1920) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these four specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens

badachschana Alberti, 1939

Original combination

Hesperia badachschana nov. sp.” Alberti, 1939 Ent. Rdsch. 56: 107.

Current combination

Pyrgus badachschana (Alberti, 1939).

Current status

Valid species.

Type material

Holotype 1♂ (ZMH 827639) (Fig. 169). “HOLOTYPE ♂ / Hesperia / badachschana” // “Badachschan / Seba[..]-Tal / Alpenwiesenzone / 2800–3000 m Mitte Juni / leg. H.&E. Kotzsch” // “Coll. Bytinski-Salz Coll / Eing. Nr. 20, 1960 Eing” // “ZMH 827639”.

Type locality

Afghanistan: Badakhshan.

Remarks

We follow SEL declaration on gender agreement (Sommerer 2002), as a result it should stay as badachschana.

picena Verity, 1920

Original combination

Hesperia foulquieri, Obth., race picena, mihi” Verity, 1920 Ent. Rec. 32: 4.

Current combination

Pyrgus foulquieri picena (Verity, 1920).

Current status

Valid subspecies.

Original material

Labelled as “Type” 4♂♂ (ZMH 833506–833509) (Fig. 170). “Picena Vrty” // “Marche / Bolognola / 2 VII 1937 / Querci” // [blank label] // “ZMH 833506”; “Picena Vrty” // “Marche / Bolognola / 8 Ag 1937 / Querci” // [blank label] // “ZMH 833507”; “Picena Vrty” // “Marche / Bolognola / 8 Ag 1937 / Querci” // [blank label] // “ZMH 833508”; “Picena Vrty” // “Marche / Bolognola / 12 Ag 1937 / Querci” // [blank label] // “ZMH 833509”.

Original locality

Italy: Bolognola, Sibillini Mts.

Remarks

Verity (1920) proposed this name as a “race” of H. foulquieri Oberthür, 1910. As it was given in trinomy and the content of the description clearly indicates that a geographical race was meant, this name is available as subspecific (article 45.6 ICZN 1999). The year on the specimen labels (1937) indicate specimens were collected 17 years after description (1920) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these four specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.

ab. alboradiata Bubacek, 1926

Original combination

Hesperia sidae Esp. var. onopordi H.-S. ab. nova alboradiata Bub.” Bubacek, 1926 Verh. Zool. Bot. Ges. Wien 76: 174.

Current combination

Pyrgus sidae ab. alboradiata (Bubacek, 1926).

Current status

Infrasubspecific and hence unavailable name.

Original material

Labelled as “Type” 1♂ (ZMH 833497) (Fig. 171). “Type / H. sidae v. onopordi / ab. alboradiata Bub. / Z. B. G. 1926 O. Bubacek” // “Italia / Alassio – Liguria / 7/6 1925 / Coll. O. Bubacek” // [blank label] // “ZMH 833497”.

Original locality

Italy: “Alassio, Liguria”.

Remarks

Bubacek (1926) proposed this name as an aberration of H. sidae v. onopordi “H.-S.” (sic, recte Rambur, 1839), therefore, as stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable. P. onopordi Rambur is a distinct species from P. sidae Esper, and both species occur sympatrically in Liguria.

Plate 1.

1. Parnassius apollo tridentina Dannehl, 1925; 2. Parnassius apollo mendolensis Dannehl, 1925; 3. Parnassius apollo sevensis Kesenheimer, 1921; 4. Parnassius apollo kricheldorffi Eisner, 1928; 5. Parnassius apollo valesiacus ab. praetexta Gelpke, 1929; 6. Parnassius apollo ab. uniformis Stauder, 1922. a. Dorsal view, b. ventral view.

Plate 2.

7. Parnassius apollo pumilus f. bispupillata Turati, 1918; 8. Parnssius apollo pumilus f. appendiculata Turati, 1918; 9. Parnassius apollo ab. lacrimans Marschner, 1911; 10. Parnassius apollo oenipontanus Stauder & Felkel, 1921; 11. Parnassius apollo serpentinicus Mayer, 1925 12. Parnassius apollo ab. marschneri Bryk, 1911. a. Dorsal view, b. ventral view.

Plate 3.

13. Parnassius apollo wenzeli Bryk, 1921; 14. Parnassius nomion gabrieli Bryk, 1934; 15. Parnassius nomion chingamensis Bryk & Eisner, 1932; 16. Parnassius nomion tsinlingensis Bryk & Eisner, 1932; 17. Parnassius nomion nominulus Staudinger, 1894; 18. Parnassius szechenyii frivaldszkyi Bang-Haas, 1928. a. Dorsal view, b. ventral view.

Plate 4.

19. Parnassius autocrator ab. privatus Kotzsch, 1936; 20. Parnassius orleans lictor Bryk & Eisner, 1937; 21. Parnassius jacquemontii buddha Bryk & Eisner, 1937; 22. Parnassius bremeri amgunensis ab. tripicta Sheljuzhko, 1928; 23. Parnassius actius melaniticus Bang-Haas, 1915; 24. Parnassius charltonius wernickei Kotzsch, 1936. a. Dorsal view, b. ventral view.

Plate 5.

25. Parnassius inopinatus Kotzsch, 1940; 26. Parnassius imperator gigas Kotzsch, 1932; 27. Parnassius mnemosyne clorinda Kolar & Querci, 1937; 28. Parnassius mnemosyne rjabovi Sheljuzhko, 1935; 29. Parnassius mnemosyne comitis Bryk, 1912; 30. Parnassius mnemosyne gigantea Staudinger, 1886. a. Dorsal view, b. ventral view.

Plate 6.

31. Parnassius epaphus beicki Bryk & Eisner, 1932; 32. Parnassius epaphus ab. rubropicta Bang-Haas, 1915; 33. Parnassius delphius candidatus Bang-Haas, 1915; 34. Parnassius jacobsoni eva Kotzsch, 1936; 35. Parnassius mnemosyne hartmanni ab. minor Frank; 36. Parnassius mnemosyne hartmanni ab. nigra Frank; 37. Parnassius mnemosyne hartmanni ab. punctata Frank; 38. Parnassius mnemosyne hartmanni ab. nigerrima Frank. a. Dorsal view, b. ventral view.

Plate 7.

39. Parnassius apollo ab. subtusalbipupillata; 40. Parnassius apollo ab. analifasciata Gelpke; 41. Papilio xuthus f. unimaculata Warnecke, 1924; 42. Battus polydamas peruanus (Fuchs, 1954); 43. Iphiclides podalirius ab. ornatissima (Warnecke, 1924); 44. Iphiclides feisthamelii f. ornatus ab. indistinctus Meinhard. a. Dorsal view, b. ventral view.

Plate 8.

45. Colias heos ab. crocopepla Warnecke, 1912; 46. Colias heos ab. rhododactyla Warnecke, 1912; 47. Colias heos ab. theia Warnecke, 1912; 48. Colias caucasica balcanica f. rebeli Schawerda, 1906; 49. Colias cocandica tatarica Bang-Haas, 1915; 50. Colias wiskotti aurea Kotzsch, 1936. a. Dorsal view, b. ventral view.

Plate 9.

51. Eurema zelia andina (Fuchs, 1928); 52. Leptidea duponcheli lorkovici Pfeiffer, 1932; 53. Leptidea sinapis f. fragilis Dannehl, 1925; 54. Leptidea sinapis ab. simbruina Dannehl, 1927; 55. Leptidea sinapis stabiarum Staudinger, 1914; 56. Anthocharis euphenoides italorum Dannehl, 1929; 57. Euchloe ausonia paravicinii Stauder, 1915; 58. Colotis evagore nouna f. flavide apicata (Stauder, 1922). a. Dorsal view, b. ventral view.

Plate 10.

59. Colotis evagore nouna f. vernalis alticola auresiaca (Stauder, 1913); 60. Colotis evagore nouna f. biformata (Stauder, 1913); 61. Colotis evagore nouna f. aestiva pyroleuca (Stauder, 1913); 62. Colotis evagore nouna ab. evagorides (Stauder, 1913); 63. Pieris brassicae italorum Stauder, 1921; 64. Pieris ergane elbursina Bytinski-Salz & Brandt, 1937; 65. Pieris ergane r. exigua Verity, 1923. a. Dorsal view, b. ventral view.

Plate 11.

66. Pieris rapae f. rossioides Stauder, 1921; 67. Pieris napi v. napaeae ab. gottingensis Meinhard; 68. Pieris brassicae ab. nigromarginata Meinhard; 69. Cigaritis acamas dueldueli Pfeiffer, 1932; 70. Lycaena alciphron f. emarginata (Warnecke, 1942); 71. Lycaena virgaureae ab. alba (Schoenfeld, 1924); 72. Lycaena virgaureae f. multipunctata (Warnecke, 1942). a. Dorsal view, b. ventral view.

Plate 12.

73. Lycaena virgaureae emilianus f. quercii (Turati, 1923); 74. Agriades pheretiades andarabi (Forster, 1938); 75. Aricia anteros f. altera (Züllich, 1929); 76. Aricia anteros r. modicior Verity, 1938; 77. Plebejus idas uliginosa (Dannehl, 1921); 78. Plebejus argyrognomon balcanica (Züllich, 1929); 79. Eumedonia eumedon mayencis Eitschberger & Steiniger, 1975; 80. Eumedonia eumedon moenus Eitschberger & Steiniger, 1975; 81. Freyeria trochylus persa (Bytinski-Salz & Brandt, 1937); 82. Freyeria trochylus persa ab. pauper (Bytinski-Salz & Brandt, 1937); 83. Glaucopsyche alexis parvandereggi Verity, 1938; 84. Lysandra coridon sibyllina (Verity, 1916). a. Dorsal view, b. ventral view.

Plate 13.

85. Phengaris alcon f. lunulata (Warnecke, 1942); 86. Phengaris arionides f. selzeri (Warnecke, 1924); 87. Phengaris euphemus peninsulae (Verity, 1923); 88. Cupido argiades ab. depuncta (Hirschke, 1903); 89. Polyommatus (Agrodiaetus) damon ausonia (Verity, 1913); 90. Polyommatus (Agrodiaetus) pfeifferi (Brandt, 1938); 91. Polyommatus (Agrodiaetus) alcestis (Zerny, 1932); 92. Polyommatus (Agrodiaetus) ardschira (Brandt, 1938); 93. Polyommatus daphnis macra f. squalida Verity, 1920; 94. Polyommatus dorylas r. correpta (Verity, 1920). a. Dorsal view, b. ventral view.

Plate 14.

95. Polyommatus escheri olympena (Verity, 1936); 96. Polyommatus icarus ab. privata (Schönfeld, 1924); 97. Polyommatus (Sublysandra) nuksani (Forster, 1937); 98. Tarucus balkanica ab. clorinda Verity, 1938; 99. Turanana panagaea ahasveros (Bytinski-Salz & Brandt, 1937); 100. Satyrium ilicis syra (Pfeiffer, 1932); 101. Satyrium spini brevicaudis (Vorbrodt, 1911); 102. Lycaena cyllarus v. romana Hartig; 103. Lysandra coridon ab. albipunctata Hartig. a. Dorsal view, b. ventral view.

Plate 15.

104. Danaus gilippus icensis (Fuchs, 1954); 105. Danaus gilippus roedingeri (Fuchs, 1954); 106. Fabriciana niobe altonevadensis (Reisser, 1927); 107. Argynnis pandora cyrnea Schawerda, 1926; 108. Argynnis pandora argentifasciata Kotzsch, 1937; 109. Euphydryas maturna f. urbanoides (Warnecke, 1942). a. Dorsal view, b. ventral view.

Plate 16.

110. Melitaea athalia tenuicula Verity, 1919; 111. Melitaea varia varissima Verity, 1914; 112. Melitaea cinxia riloensis Züllich, 1936; 113. Melitaea deione ab. mediofasciata Bubacek, 1926; 114. Melitaea diamina ab. johni Stauder, 1922; 115. Melitaea trivia balcanica Züllich, 1936; 116. Melitaea trivia catapelioides Stauder, 1918; 117. Melitaea trivia ignasiti de Sagarra, 1926. a. Dorsal view, b. ventral view.

Plate 17.

118. Aglais urticae guhni (Tschauner, 1926); 119. Araschnia levana f. clarior Warnecke, 1959; 120. Araschnia levana prorsa ab. marginelineata Horch, 1932; 121. Arethusana arethusa alphea (Warnecke, 1919); 122. Brintesia circe asperomontana (Stauder, 1921). a. Dorsal view, b. ventral view.

Plate 18.

123. Coenonympha arcania f. rufobrunnea Warnecke, 1942; 124. Coenonympha arcania tyrrhena Stauder, 1915; 125. Coenonympha corinna ab. energica Bubacek, 1923; 126. Coenonympha corinna ab. caeca Bubacek, 1923; 127. Coenonympha corinna ab. anophthalmica Bubacek, 1923; 128. Coenonympha corinna ab. saturata Bubacek, 1923; 129. Coenonympha corinna ab. nigricans Bubacek, 1923. a. Dorsal view, b. ventral view.

Plate 19.

130. Coenonympha corinna ab. macrophthalmica Bubacek, 1923; 131. Coenonympha corinna ab. gynandra Bubacek, 1923; 132. Coenonympha glycerion f. pulcherrima Warnecke, 1942; 133. Coenonympha glycerion ab. oikeia Dannehl, 1927; 134. Coenonympha leander iranica Schwingenschuss, 1939; 135. Coenonympha oedippus rhenana Gradl, 1933. a. Dorsal view, b. ventral view.

Plate 20.

136. Coenonympha pamphilus galvagnii Stauder, 1923; 137. Coenonympha tullia f. maculata Warnecke, 1942; 138. Erebia flavofasciata Heyne, 1895; 139. Erebia aethiops peneplana Berger, 1936; 140. Erebia epiphron clorinda Hartig, 1940; 141. Erebia epiphron vogesiaca Goltz, 1914. a. Dorsal view, b. ventral view.

Plate 21.

142. Erebia melas leonhardi Fruhstorfer, 1918; 143. Erebia meolans paradisi Dannehl, 1929; 144. Erebia ottomana benacensis Warren, 1933; 145. Paralasa kotzschae (Goltz, 1937); 146. Hipparchia autonoe maxima (Bang-Haas, 1933). a. Dorsal view, b. ventral view.

Plate 22.

147. Hipparchia fatua persiscana (Verity, 1937); 148. Hipparchia neomiris ab. pallida (Bubacek, 1923); 149. Hipparchia semele ab. albinervata (Warnecke, 1944); 150. Lymanopoda shefteli siviae Fuchs, 1954; 151. Melanargia arge cocuzzana Stauder, 1914. a. Dorsal view, b. ventral view.

Plate 23.

152. Lasiommata petropolitana ominata ab. huenei (Hirschke, 1910); 153. Lasiommata felix (Warnecke, 1929); 154. Pararge aegeria ab. diluta Bubacek, 1923; 155. Pararge aegeria egerides f. albiplaga Warnecke, 1942; 156. Lasiommata megera lyssa ab. hermini (Hirschke, 1910); 157. Punargentus angusta picota (Fuchs, 1954). a. Dorsal view, b. ventral view.

Plate 24.

158. Pyronia cecilia arminii (Stauder, 1916); 159. Satyrus ferula liupiuschani Bang-Haas, 1933; 160. Coenonympha phryne kintschouensis (Bang-Haas, 1939); 161. Melanargia galathea v. sakaria ab. minor Meinhard; 162. Epinephele ida ab. caeca Dannehl; 163. Satyrus actaea achaltekkensis Holik. a. Dorsal view, b. ventral view.

Plate 25.

164. Satyrus actaea rossarmenia Bang-Haas; 165. Hamearis lucina ab. friesi Meinhard; 166. Carcharodus alceae wissmanni Warnecke, 1934; 167. Spialia orbifer orbifer v. postesseloides Verity, 1938; 168. Pyrgus alveus centralitaliae (Verity, 1920); 169. Pyrgus badachschana (Alberti, 1939); 170. Pyrgus foulquieri picena (Verity, 1920); 171. Pyrgus sidae ab. alboradiata (Bubacek, 1926). a. Dorsal view, b. ventral view.

Discussion

With more than 2,000 type specimens of butterflies and moths (Lepidoptera), the Zoological Museum of Hamburg (ZMH) is one of the most important natural history archives in Germany. In this catalogue, a total of 414 specimens labelled as “types” belonging to the superfamily Papilionoidea are listed and discussed. This includes 23 specimens of 16 “in litteris” (= unavailable) names, 43 topotypes (non-type specimens), 97 original specimens of invalid infrasubspecific entities under the ICZN code, and 251 type specimens (Fig. 172; Suppl. material 2: Table S2). Out of 251 type specimens, 18 specimens represent species (nine valid species), and 233 specimens are subspecies (representing 74 subspecies: 44 valid names and 30 synonyms) (Suppl. material 1: Table S1, Suppl. material 2: S2; Fig. 172). Of these 414 specimens, 171 are primary types (8 holotypes/lectotypes and 163 syntypes), 80 are secondary types, 120 are considered of invalid infrasubspecific rank (original specimens), and 43 specimens are re-labelled as “Non-type” specimens (topotype) (Fig. 172).

The comparison of our updated catalogue to the type catalogue of Weidner (1974) reveals 95 new additions (237 specimens) to the type collection and type material of one hersperiid species (i.e., Adopaea pfeifferi Bytinski-Salz & Brandt, 1937) being potentially lost (one specimen) (Suppl. material 1: Tables S1, Suppl. material 2: Table S2). The latter species is currently a junior subjective synonym of Thymelicus hyrax (Lederer, 1861). New additions to the type collection since the Weidner catalogue (1974) do not necessarily reflect “new type additions” in all instances. Except for a few “type” specimens donated after 1974 to the ZMH collection (e.g., from the former collection of the Altona museum), many specimens were already during Weidner’s time in the collection, but were either misidentified or misplaced. Details of the changes and updates including the new types, changes to the type status, and changes to the specimen counts since the last catalogue of Weidner (1974) were summarized in Suppl. material 1: Tables S1, Suppl. material 2: S2.

Five countries (Italy – 37, Germany – 25, China – 16, France – 13, and Afghanistan – 10) represent the majority of type localities for 60% (101 of 172 “names”) of the Papilionoidea type collection followed by Iran and Austria (each represented by 8 “type” localities), Russia, Greece, Peru and Algeria (each with 6 “type” localities) (Fig. 173). Most type specimens come from the collection of Dr. George Warnecke, mainly from Europe and the Middle East (52 taxa), followed by the collection of Dr. Hanan Bytinski-Salz (20 taxa). The oldest type specimen of Papilionoidea at ZMH is the lectotype of Colias aurorina ab. crocopepla Warnecke, 1912 collected in Amur in 1887. The oldest described types are Parnassius mnemosyne giganteus Staudinger, 1886 and Parnassius nomion nominulus Staudinger, 1894, both are valid subspecies names from the collections of H. Rödinger and G. Warnecke, respectively, followed by Erebia flavofasciata Heyne, 1895, a valid species from the collection of von Nolte.

Figure 172.

Type status among 414 “type” specimens of the lepidopteran superfamily Papilionoidea deposited in the Zoological Museum of Hamburg (ZMH).

Figure 173.

Geographical distribution of “type” localities among 171 representative “type” specimens of the lepidopteran superfamily Papilionoidea deposited in the Zoological Museum of Hamburg (ZMH). Category A (six type localities) includes Algeria, Greece, Peru and Russia; category B (three type localities) includes Switzerland, Turkey and Spain; category C (two type localities) includes Armenia, Croatia and Yemen; category D (one type locality) includes Turkmenistan, Sweden, Romania, Macedonia. Lebanon, Japan, Estonia, Czech Republic, Bosnia and Herzegovina, and Belgium.

Acknowledgements

This project was funded by the CeNak “Digitalisierung der CeNak-Sammlungen”. We also like to thank HASPA Lotterie Sparen for funding boxes and the City of Hamburg for further funding via Bussgeldfonds. We thank the Gesellschaft der Freunde und Förderer des Zoologischen Museums Hamburg for providing financial support to the collections. The authors also acknowledge the members of the other Zoological Collections at the CeNak for their patience during our intensive photography sessions. Comments by Vladimir Lukhtanov (St. Petersburg, Russia), Niklas Wahlberg (Lund, Sweden), Alberto Zilli (London, UK), Karen McLachlan Hamilton (Ottawa, Canada) and Danilo Harms (Hamburg, Germany) substantially improved the manuscript.

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